Krikken, J., and J. Huijbregts. “New Guinea Onthophagus: Taxonomy of Ten Small, Unicolored New Species (Coleoptera: Scarabaeidae: Scarabaeinae)”. Zootaxa 3619, no. 5 (March 4, 2013): 501–525. Accessed April 21, 2025. https://mapress.com/zt/article/view/zootaxa.3619.5.1.