Schmidt, Chris. “Molecular Phylogenetics of Ponerine Ants (Hymenoptera: Formicidae: Ponerinae)”. Zootaxa 3647, no. 2 (May 9, 2013): 201–250. Accessed April 22, 2025. https://mapress.com/zt/article/view/zootaxa.3647.2.1.