Perkins, Philip D. “A Revision of the Water Beetle Genus Hydraena Kugelann for Southern Africa (Coleoptera: Hydraenidae)”. Zootaxa 3758, no. 1 (January 29, 2014): 1–92. Accessed April 22, 2025. https://mapress.com/zt/article/view/zootaxa.3758.1.1.