Kodada, Ján, Manfred A. Jäch, and Fedor Čiampor Jr. “Ancyronyx reticulatus and A. pulcherrimus, Two New Riffle Beetle Species from Borneo, and Discussion about Elmid Plastron Structures (Coleoptera: Elmidae)”. Zootaxa 3760, no. 3 (February 3, 2014): 383–395. Accessed April 22, 2025. https://mapress.com/zt/article/view/zootaxa.3760.3.5.