Bilton, David T. “New Species and New Records of Pterosthetops: Eumadicolous Water Beetles of the South African Cape (Coleoptera, Hydraenidae)”. Zootaxa 3811, no. 4 (June 5, 2014): 438–462. Accessed January 2, 2026. https://mapress.com/zt/article/view/zootaxa.3811.4.2.