Maruyama, Munetoshi, Shûhei Yamamoto, and Taro K. Eldredge. “Synopsis of the Japanese Species of Aleocharinae (Coleoptera: Staphylinidae), With Review of the Type Specimens I. Tribes Himalusini and Leucocraspedini”. Zootaxa 3887, no. 3 (November 25, 2014): 393–400. Accessed May 1, 2025. https://mapress.com/zt/article/view/zootaxa.3887.3.6.