Duellman, William E., and S. Blair Hedges. “Two New Minute Species of Phrynopus (Lissamphibia: Anura) from the Cordillera Oriental in Peru”. Zootaxa 1675, no. 1 (January 9, 2008): 59–66. Accessed April 21, 2025. https://mapress.com/zt/article/view/zootaxa.1675.1.4.