Edmonds, W. D. “A New Species of Coprophanaeus Olsoufieff (Coleoptera: Scarabaeidae) from Bolivia”. Zootaxa 1723, no. 1 (March 10, 2008): 42–46. Accessed April 20, 2025. https://mapress.com/zt/article/view/zootaxa.1723.1.2.