Japoshvili, George, and Eduard Khachikov. “List of Staphylinids of Lagodekhi Reserve With Some New Records from Sakartvelo (Georgia)”. Zootaxa 4767, no. 3 (April 28, 2020): 495–500. Accessed April 22, 2025. https://mapress.com/zt/article/view/zootaxa.4767.3.9.