Chaparro, Juan C., Jose M. Padial, and Ignacio de La Riva. “Two Sympatric New Species of Phrynopus (Anura: Strabomantidae) from Yanachaga Chemillen National Park (central Peruvian Andes)”. Zootaxa 1761, no. 1 (May 2, 2008): 49–58. Accessed April 21, 2025. https://mapress.com/zt/article/view/zootaxa.1761.1.5.