Andújar, Carmelo, and Vasily V. Grebennikov. “Endogean Beetles (Coleoptera) of Madagascar: Deep Soil Sampling and Illustrated Overview”. Zootaxa 4963, no. 2 (April 19, 2021): 317–334. Accessed April 3, 2025. https://mapress.com/zt/article/view/zootaxa.4963.2.4.