Penz, Carla M. “The Phylogeny of Opoptera Butterflies, and an Assessment of the Systematic Position of O. staudingeri (Lepidoptera, Nymphalidae)”. Zootaxa 1985, no. 1 (January 21, 2009): 1–20. Accessed April 7, 2025. https://mapress.com/zt/article/view/zootaxa.1985.1.1.