Tian, Ming-Yi, and Thierry Deuve. “A Review of the Genus Actenoncus Chaudoir (Coleoptera: Caraboidea: Orthogoniini)”. Zootaxa 2135, no. 1 (June 18, 2009): 57–64. Accessed April 20, 2025. https://mapress.com/zt/article/view/zootaxa.2135.1.2.