Smith, Derek J., and Steve Shattuck. “Six New, Unusually Small Ants of the Genus Leptomyrmex (Hymenoptera: Formicidae)”. Zootaxa 2142, no. 1 (June 26, 2009): 57–68. Accessed April 19, 2025. https://mapress.com/zt/article/view/zootaxa.2142.1.5.