Yoshimura, Masashi, and Brian L. Fisher. “A Revision of Male Ants of the Malagasy Region (Hymenoptera: Formicidae): Key to Genera of the Subfamily Proceratiinae”. Zootaxa 2216, no. 1 (September 1, 2009): 1–21. Accessed April 20, 2025. https://mapress.com/zt/article/view/zootaxa.2216.1.1.