Perkins, Philip D. “A Revision of the Australian Endemic Water Beetle Genus Gymnanthelius Perkins (Coleoptera: Hydraenidae)”. Zootaxa 585, no. 1 (July 27, 2004): 1–39. Accessed April 11, 2025. https://mapress.com/zt/article/view/zootaxa.585.1.1.