Queney, Pierre. “Two New Species of Berosus Leach from Brazil (Coleoptera: Hydrophilidae: Berosini)”. Zootaxa 2506, no. 1 (June 15, 2010): 51–58. Accessed April 20, 2025. https://mapress.com/zt/article/view/zootaxa.2506.1.4.