Muilwijk, Jan, Boris M. Kataev, and David W. Wrase. “Review of the Subgenus Lyrothorax of the Genus Pterostichus (Coleoptera: Carabidae) in the Western Palaearctic, With Descriptions of Two New Species”. Zootaxa 5468, no. 2 (June 12, 2024): 255–274. Accessed April 26, 2025. https://mapress.com/zt/article/view/zootaxa.5468.2.2.