Génier, François. “Three New Species of Onthophagus Latreille, 1802 from Bioko Island, Equatorial Guinea, Belonging to d’Orbigny’s Group IX, With a Taxonomic Note on O. denudatus D’Orbigny, 1902 (Coleoptera: Scarabaeidae: Scarabaeinae: Onthophagini)”. Zootaxa 5831, no. 3 (June 16, 2026): 413–426. Accessed June 17, 2026. https://mapress.com/zt/article/view/zootaxa.5831.3.6.