Dehling, J. Maximilian. “A New Karst-Dwelling Species of Kalophrynus (Anura: Microhylidae) from Gunung Mulu National Park, Borneo, Malaysia”. Zootaxa 2737, no. 1 (January 12, 2011): 49–60. Accessed June 9, 2025. https://mapress.com/zt/article/view/zootaxa.2737.1.4.