Medina CA, Molano F, Scholtz CH. Morphology and terminology of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) male genitalia. Zootaxa [Internet]. 2013 Mar. 15 [cited 2025 Apr. 21];3626(4):455–476. Available from: https://mapress.com/zt/article/view/zootaxa.3626.4.3