Cupello M, Vaz-De-Mello FZ. Taxonomic revision of the South American dung beetle genus Gromphas Brullé, 1837 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini: Gromphadina). Zootaxa [Internet]. 2013 Oct. 24 [cited 2025 Apr. 24];3722(4):439–482. Available from: https://mapress.com/zt/article/view/zootaxa.3722.4.2