Brown, J.L., Twomey, E., Amézquita, A., de Souza, M.B., Caldwell, J.P., Lötters, S., Von May, R., Melo-Sampaio, P.R., Mejía-Vargas, D., Perez-Peña, P., Pepper, M., Poelman, E.H., Sanchez-Rodriguez, M. & Summers, K. (2011) A taxonomic revision of the Neotropical poison frog genus Ranitomeya (Amphibia: Dendrobatidae). Zootaxa, 3083 (1), 1–120.