Gareca, Y. & Blandin, P. (2011) Morpho (Morpho) helenor (Cramer) (Lepidoptera, Nymphalidae, Morphinae) in Bolivia: Geographical distribution and ecological plasticity, with a description of a new subspecies. Zootaxa, 3130 (1), 30–56.
https://doi.org/10.11646/zootaxa.3130.1.2