Johnson, C.B., S. H., E.Q., Anuar, S., Muin, M.A., Wood, Jr., P.L., Grismer, J.L., Greer, L.F., Onn, C.K., Ahmad, N., Bauer, A.M. & Grismer, L.L. (2012) Phylogeography, geographic variation, and taxonomy of the Bent-toed Gecko Cyrtodactylus quadrivirgatus Taylor, 1962 from Peninsular Malaysia with the description of a new swamp dwelling species. Zootaxa, 3406 (1), 39–58.
https://doi.org/10.11646/zootaxa.3406.1.3