Hernández-Roldán, J.L., Munguira, M.L., Wagner, W. & Vila, R. (2012) Comparative analysis and taxonomic use of the morphology of immature stages and natural history traits in European species of Pyrgus Hübner (Lepidoptera: Hesperiidae, Pyrginae). Zootaxa, 3470 (1), 1–71.
https://doi.org/10.11646/zootaxa.3470.1.1