Sasakawa, K. & Kubota, K. (2006) Nebria tenuicaulis sp. nov., a sympatric species with Nebria sadona ohdaiensis Nakane, with studies on the phylogeny of the subgenus Sadonebria Ledoux & Roux (Coleoptera: Carabidae). Zootaxa, 1306 (1), 41–50.