Howden, H.F. (2006)

New species and a new subgenus of South American Neoathyreus Howden and Martínez (Coleoptera: Geotrupidae: Athyreini)

. Zootaxa, 1137 (1), 37–52.
https://doi.org/10.11646/zootaxa.1137.1.2