Borges, A., de Sousa, L. & Manzanilla, J. (2006)

Description of a new Tityus species (Scorpiones: Buthidae) from Sierra de Portuguesa, western Venezuela, based on morphological and mitochondrial DNA evidence

. Zootaxa, 1107 (1), 49–68.
https://doi.org/10.11646/zootaxa.1107.1.2