Venegas, P.J., Barboza, A.C., Riva, I.D.L. & Padial, J.M. (2018)

A new species of Phrynopus from the northeastern Andes of Peru, its phylogenetic position, and notes on the relationships of Holoadeninae (Anura: Craugastoridae)

. Zootaxa, 4446 (4), 501–524.