Kohlmann, B., Solís, Ángel, Elle, O., Soto, X. & Russo, R. (2007)

Biodiversity, conservation, and hotspot atlas of Costa Rica:  a dung beetle perspective (Coleoptera: Scarabaeidae: Scarabaeinae)

. Zootaxa, 1457 (1), 1–34.
https://doi.org/10.11646/zootaxa.1457.1.1