Rueda-Almonacid, J.V., Rada, M., Sánchez-Pacheco, S.J., Velásquez-Álvarez, Álvaro A. & Quevedo, A. (2006)

Two new and exceptional poison dart frogs of the genus Dendrobates (Anura: Dendrobatidae) from the Northeastern Flank of the cordillera Central of Colombia

. Zootaxa, 1259 (1), 39–54.