Pfingstl, T., Schäffer, S., Ebermann, E. & Krisper, G. (2009) Differentiation between two epilittoral species, Scutovertex arenocolus spec. nov. and Scutovertex pilosetosus Polderman (Acari: Oribatida) from different European coasts. Zootaxa, 2153 (1), 35–54.
https://doi.org/10.11646/zootaxa.2153.1.3