Fukuda, Y., Ogawa, R. & Hori, M. (2019) The reclassification of Sophiodela and other tiger beetles (Coleoptera, Cicindelidae) based on the structure of the everted internal sac of the male genitalia. Zootaxa, 4661 (2), 271–308.
https://doi.org/10.11646/zootaxa.4661.2.3