Magalhães, C., Wehrtmann, I.S., Lara, L.R. & Mantelatto, F.L. (2015)

Taxonomy of the freshwater crabs of Costa Rica, with a revision of the genus Ptychophallus Smalley, 1964 (Crustacea: Decapoda: Pseudothelphusidae)

. Zootaxa, 3905 (3), 301–344.