Marinho, M.M.F., Carvalho, F.R., Langeani, F. & Tatsumi, F.L. (2008) A new Hemigrammus Gill from upper rio Parana system, Southeastern Brazil (Characiformes: Characidae). Zootaxa, 1724 (1), 52–60.
https://doi.org/10.11646/zootaxa.1724.1.2