Ahmed, Q., Thandar, A.S. & Ali, Q.M. (2020)

Holothuria (Lessonothuria) insignis Ludwig, 1875 (formally resurrected from synonymy of H. pardalis Selenka, 1867) and Holothuria (Lessonothuria) lineata Ludwig, 1875—new additions to the sea cucumber fauna of Pakistan, with a key to the subgenus Lessonothuria Deichmann (Echinodermata: Holothuroidea)

. Zootaxa, 4767 (2), 307–318.