Vigle, G.O., Coloma, L.A., Santos, J.C., Hernandez-Nieto, S., Ortega-Andrade, H.M., Paluh, D.J. & Read, M. (2020)

A new species of Leucostethus (Anura: Dendrobatidae) from the Cordillera Mache-Chindul in northwestern Ecuador, with comments on similar Colostethus and Hyloxalus

. Zootaxa, 4896 (3), 342–372.
https://doi.org/10.11646/zootaxa.4896.3.2