Gunter, N.L. & Weir, T.A. (2021)

Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): review of the L. ustulatus, L. storeyi, and L. nitidus species groups and description of eight new species

. Zootaxa, 4923 (1), 1–66.