Tojo, K., Miyairi, K., Kato, Y., Sakano, A. & Suzuki, T. (2021)

A description of the second species of the genus Bleptus Eaton, 1885 (Ephemeroptera: Heptageniidae) from Japan, and phylogenetic relationships of two Bleptus mayflies inferred from mitochondrial and nuclear gene sequences

. Zootaxa, 4974 (2), 333–360.
https://doi.org/10.11646/zootaxa.4974.2.5