Ogden, T.H., Osborne, J.T., Jacobus, L.M. & Whiting, M.F. (2009) Combined molecular and morphological phylogeny of Ephemerellinae (Ephemerellidae: Ephemeroptera), with remarks about classification. Zootaxa, 1991 (1), 28–42.
https://doi.org/10.11646/zootaxa.1991.1.2