Kohlmann, B. & Solís, Ángel (2025) A review of the species groups of the Western Hemisphere Onthophagus Latreille (Coleoptera: Scarabaeidae: Scarabaeinae) using COI barcoding and gene trees. Zootaxa, 5604 (4), 401–447.
https://doi.org/10.11646/zootaxa.5604.4.1