Fresneda, J., Ortuño, V.M., Arribas, Óscar & Salgado, J.M. (2026) Leonesiella tetratriches sp. nov. from the Cantabrian Mountains (León, Spain)—first species of Cantabrian Leptodirini found in mesovoid shallow substratum (MSS) (Coleoptera, Leiodidae, Cholevinae, Leptodirini). Zootaxa, 5757 (3), 261–270.