Coscarón, S., Esquivel, D.R.M., Moulton, J.K., Coscarónarias, C.L. & Bernal, S.I. (2004) Simulium (Hearlea) Vargas, Mart nez Palacios, & D az N jera (Diptera: Simuliidae): Taxonomic revision and cladistic analysis. Zootaxa, 396 (1), 1–52.