Skip to main content Skip to main navigation menu Skip to site footer
Type: Article
Published: 2026-01-06
Page range: 114–175
Abstract views: 49
PDF downloaded: 38

New taxa of Mesophotic Asteroidea from Ashmore Reef, Western Australia

Dept. of Invertebrate Zoology, National Museum of Natural History, Smithsonian Institution, Washington, DC. 20560
Collections and Research, Western Australian Museum, Perth, Western Australia, 6106
Collections and Research, Western Australian Museum, Perth, Western Australia, 6106, School of Biological Sciences, University of Western Australia, Crawley, Western Australia, 6009
Collections and Research, Western Australian Museum, Perth, Western Australia, 6106, School of Molecular and Life Sciences, Curtin University, Bentley, Western Australia, 6012
Mesophotic Ecosystems MCE Goniasteridae Pterasteridae Podosphaerasteridae Ophidiasteridae Asterinidae Brisingaster Western Australia Indian Ocean associates Polynoidae Eulimidae

Abstract

We report on 27 asteroid specimens collected by the RV Falkor, using the ROV SuBastian, from mesophotic depths on Ashmore Reef in Western Australia, 2021. Of the total, 10 new species were collected, 9 of which are described here. Sixteen species were new occurrences for Western Australia. One additional shallow water species was observed at mesophotic depths but not collected. Each collected specimen was identified as a different species. In the Valvatida, this included Goniasteridae (n=9), Mithrodiidae (n=1), Ophidiasteridae (n=4), Asterinidae (n=3), Asterodiscididae (n=2), Oreasteridae (n=2), Podosphaerasteridae (n=1), and the Echinasteridae (n=3). Also present were two non-valvatidan groups, a brisingid from the Forcipulatacea and a new species of pterasterid in the genus Euretaster from the Velatida. Consistent with other accounts of mesophotic asteroid distribution, some of the species observed here, such as the goniasterid Churaumiastra hoshi Mah et al. 2024 have been found to be widely distributed across the Indo-Pacific.

References

  1. Antokhina, T.I. & Britayev, T.A. (2012) Sea stars and their macrosymbionts in the Bay of Nhatrang, Southern Vietnam. Paleontological Journal, 46, 894–908. https://doi.org/10.1134/S0031030112080023
    Arai, M. & Fujita, T. (2021) Sea Stars of families Ophidiasteridae and Goniasteridae (Echinodermata: Asteroidea) from the Mesophotic Zone of the Ogasawara Islands, including two new species. Species Diversity, 26 (1), 7–21. https://doi.org/10.12782/specdiv.26.7
    Aziz, A. (1986) La faune d’asterides (Ecinodermata) de la region indo-malaise: taxonomie, zoogéographie et bathymétrie. Marine Biology Laboratory, Université libre de Bruxelles, Brussels, 598 pp.
    Aziz, A. & Jangoux, M. (1985) On the status and affinities of the goniasterid genus Calliaster Gray, 1840. In: Keegan, B.F. & O’Connor, B.D.S. (Eds.), Proceedings of the 5th International Echinoderm Conference, Galway, 1984, pp. 585–586. https://doi.org/10.1201/9781003079224-116
    Baker, A.N. & Marsh, L.M. (1976) The rediscover of Halityle regularis Fisher (Echinodermata: Asteroidea). Records of the Western Australian Museum, 4 (2), 107–116.
    Baker, E., Puglise, K.A. & Harris, P.T. (2016) Mesophotic coral ecosystems: A lifeboat for coral reefs? UN Environment, GRID-Arendal, Norway. Available from: https://policycommons.net/artifacts/2390329/mesophotic-coral-ecosystems/3411646/ (accessed 1 June 2023) [CID: 20.500.12592/mq8sjt]
    Bell, F.J. (1884) Contributions to the Systematic Arrangement of the Asteroidea.-II. The Species of Oreaster. Proceedings of the Zoological Society of London, 52 (1), 57–87. https://doi.org/10.1111/j.1096-3642.1884.tb02810.x
    Bernasconi, I. (1937) Asteroideos argentinos.1. Familia Pterasteridae. Anales del Museo argentino de Ciencias Naturales Bernardino Rivadavia, Buenos Aires, 39, 167–187.
    Birkeland, C. & Lucas, J.S. (1990) Acanthaster planci: Major Management Problem of Coral Reefs. CRC Press, Boca Raton, Florida, 257 pp.
    Blake, D.B. (1980) On the affinites of three small sea‐star families. Journal of Natural History, 14, 163–182. https://doi.org/10.1080/00222938000770171
    Bongaerts, P. & Smith, T.B. (2019) Beyond the “Deep Reef Refuge” hypothesis: a conceptual framework to characterize persistence at depth. In: Loya, Y., Puglise, K.A. & Bridge, T.C.L. (Eds.), Mesophotic Coral Ecosystems. Coral Reffs of the World. Vol. 12. Springer, Cham, pp. 881–895. https://doi.org/10.1007/978-3-319-92735-0_45
    Bongaerts, P., Cooke, I.R., Ying, H., Wels, D., den Haan, S., Hernandez-Agreda, A., Brunner, C.A., Dove, S., Englebert, N., Eyal, G., Forêt, S., Grinblat, M., Hay, K.B., Harii, S., Hayward, D.C., Lin, Y., Mihaljević, M., Moya, A., Muir, P., Sinniger, F., Smallhorn-West, P., Torda, G., Ragan, M.A., van Oppen, M.J.H., & Hoegh-Guldberg, O. (2021) Morphological stasis masks ecologically divergent coral species on tropical reefs. Current Biology, 31 (e8), 2286–2298. https://doi.org/10.1016/j.cub.2021.03.028
    Branch, G.M., Griffiths, C.L., Branch, M.L. & Beckley, L.E. (2010) Two Oceans: A guide to the marine life of southern Africa. Struik Nature, Cape Town, 456 pp.
    Bridge, T.C., Done, T.J., Friedman, A., Beaman, R.J., Williams, S.B., Pizarro, O. & Webster, J.M. (2011) Variability in mesophotic coral reef communities along the Great Barrier Reef, Australia. Marine Ecology Progress Series, 428, 63–75. https://doi.org/10.3354/meps09046
    Bruce, A.J. (1983) The Pontoniine shrimp fauna of Australia. Australian Museum Memoir, 18, 195–218. https://doi.org/10.3853/j.0067-1967.18.1984.385
    Carmignani, A., Radice, V.Z., McMahon, K.M., Holman, A.I., Miller, K., Grice, K. & Richards, Z. (2023) Levels of autotrophy and heterotrophy in mesophotic corals near the end photic zone. Frontiers in Marine Science, 10, 1089746. https://doi.org/10.3389/fmars.2023.1089746
    Cherbonnier, G. (1970) Podosphaeraster thalassae, nov.sp., espece actuelle d’Asterie de la famille jurassique des Sphaerasteridae. Comptes Rendus Academie des Sciences, Paris, 271, 203–206.
    Clark, A.H. (1916) Seven new genera of echinoderms. Journal of the Washington Academy of Sciences, 6 (5), 115–122. https://doi.org/10.5962/bhl.part.10912
    Clark, A.H. (1952) Echinoderms from the Marshall Islands. Proceedings of the U.S. National Museum, 102, 265–303. https://doi.org/10.5479/si.00963801.102-3302.265
    Clark, A.M. & Wright, C.W. (1962) A new genus and species of recent starfishes belonging to the aberrant family Sphaerasteridae, with notes on the possible origin and affinities of the family. Annals of the Magazine of Natural History, Series 13, 5 (52), 243–251. https://doi.org/10.1080/00222936208651241
    Clark, A.M. (1967) Notes on asteroids in the British Museum (Natural History) V. Nardoa and some other ophidiasterids. Bulletin of the British Museum (Natural History), Zoology, 15 (4), 169–198. https://doi.org/10.5962/bhl.part.27517
    Clark, A.M. (1974) Notes on some echinoderms from southern Africa. Bulletin of the British Museum of Natural History (Zoology), 26 (6), 423–487, 3 pls., 16 figs., 1 map, 3 tabs. https://doi.org/10.5962/bhl.part.210
    Clark, A.M. (1989) An index of names of recent Asteroidea. Part 1. Paxillosida and Notomyotida. In: Jangoux, M. & Lawrence, J.M. (Eds.), Echinoderm Studies, 3 (2), pp. 25–347.
    Clark, A.M. (1993) An index of names of recent Asteroidea. Part 2. Valvatida. Echinoderm Studies, 4, 187–366. https://doi.org/10.1201/9781003072553-4
    Clark, A.M. (1997) An index of names of recent Asteroidea. Part 3. Velatida and Spinulosida. Echinoderm Studies, 5, 183–250. https://doi.org/10.1201/9781003077909-5
    Clark, A.M. & Mah, C. (2001) An index of names of recent Asteroidea. Part 4. Forcipulatida and Brisingida. Echinoderm Studies, 6, 229–347.
    Clark, A.M. & Courtman-Stock, J. (1976) The Echinoderms of Southern Africa. British Museum of Natural History, London, 277 pp.
    Clark, A.M. & Downey, M.E. (1992) Starfishes of the Atlantic. Chapman and Hall, London, 794 pp.
    Clark, A.M. & Rowe, F.W.E. (1971) Monograph of Shallow-water Indo-West Pacific Echinoderms. British Museum (Natural History) Publications, 690, i–ix + 1–238, 31 pls.
    Clark, A.M. & Wright, C.W. (1962) A new genus and species of recent starfishes belonging to the aberrant family Sphaerasteridae, with notes on the possible origin and affinities of the family. Annals of the Magazine of Natural History, Series 13, 5 (52), 243–251. https://doi.org/10.1080/00222936208651241
    Clark, H.E.S. & McKnight, D.G. (2000) The marine fauna of New Zealand: Echinodermata: Asteroidea (Sea-stars), Order Paxillosida. NIWA Biodiversity Memoir, 116, 1–196.
    Clark, H.E.S. & McKnight, D.G. (2001) The marine fauna of New Zealand: Echinodermata: Asteroidea (Sea-stars), Order Valvatida. NIWA Biodiversity Memoir, 117, 1–269.
    Clark, H.L. (1914) The echinoderms of the Western Australian Museum. Records of the Western Australian Museum, 1 (3), 132–173.
    Clark, H.L. (1921) The echinoderm fauna of Torres Strait: its composition and its origin. Department of Marine Biology of the Carnegie Institute, 10, i–vi + 1–223, 38 pls. https://doi.org/10.5962/bhl.title.14613
    Clark, H.L. (1926) Some sea stars from the Riksmuseum, Stockholm. Arkiv for Zoologi, 18A (8), 1–8.
    Clark, H.L. (1946) The Echinoderm fauna of Australia. Its composition and its origin. Publications of the Carnegie Institution of Washington, 566, 1–567.
    Coleman, H.L. (1911) Scientific results of the trawling expedition of H.M.C.S. Thetis off the coast of New South Wales, in February and March, 1989. Supplement to Echinodermata. Memoirs of the Australian Museum, 4 (14), 699–701, pl. 83. https://doi.org/10.3853/j.0067-1967.4.1911.1510
    Döderlein, L. (1888) Echinodermen von Ceylon. Bericht über die von den Herren Dres Sarasin gesammelten Asteroidea, Ophiuroidea und Echinoidea. Zoologische Jahrbücher, 3, 821–846. https://doi.org/10.5962/bhl.part.1933
    Döderlein, L. (1916) Uber die Gattung Oreaster und verwandte. Zoologische Jahrbücher, 40, 409–440.
    Döderlein, L. (1922) Uber die Gattung Calliaster Gray. Bijdragen tot de Dierkunde, 1922, 47–52. https://doi.org/10.1163/26660644-02201008
    Döderlein, L. (1935) Die Asteriden der Siboga-Exped. III. Oreasteridae. Siboga-Expedition, 46 (2), 71–110, pls. 15–20.
    Döderlein, L. (1936) Die Asteriden der Siboga-Exped. III. Die unterfamile Oreasterinae. Siboga-Expedition, 46 (2), 295–369, pls. 21–32.
    Domantay, J.S. & Roxas, H.A. (1938) The littoral asteroidea of Port Galera Bay and adjacent waters. Philippine Journal of Science, 65 (3), 203–237.
    Downey, M. (1968) A note on the Atlantic species of the starfish genus Linckia. Proceedings of the Biological Society of Washington, 81, 41–44.
    Downey, M.E. (1973) Starfishes from the Caribbean and the Gulf of Mexico. Smithsonian Contributions to Zoology, 126, 1–158. https://doi.org/10.5479/si.00810282.126
    Dujardin, M.F. & Hupé, M.H. (1862) Histoire naturelle des zoophytes échinodermes: comprenant la description des crinoïdes, des ophiurides, des astérides, des échinides et des holothurides. Paris: Libraire Encyclopédique de Roret, 627 pp., 10 pls. https://doi.org/10.5962/bhl.title.10122
    Eernisse, D.J. & Strathmann, M.F. (2010) A surprising species diversity in west coast Henricia (Asteroidea: Spinulosida). In: Harris, L.G., Böttger, S.A., Walker, C.W. & Lesser, M.P. (Eds.), Echinoderms: Durham. Proceedings of the 12th International Echinoderm Conference, 7–11 August 2006, Durham, New Hampshire, U.S.A. CRC Press, Boca Raton, Florida, pp. 350–353. https://doi.org/10.1201/9780203869543-8
    Fisher, W.K. (1908) Necessary changes in the nomenclature of starfishes. Smithsonian Miscellaneous Collections, Quarterly, 52, 87–93.
    Fisher, W.K. (1911) Hyalinothrix, a new genus of starfishes from the Hawaiian Islands. Proceedings of the United States National Museum, 39, 659–664. https://doi.org/10.5479/si.00963801.39-1807.659
    Fisher, W.K. (1913) New starfishes from the Philippine Islands, Celebes, and the Moluccas. Proceedings of the United States National Museum, 46, 201–222. https://doi.org/10.5479/si.00963801.46-2022.201
    Fisher, W.K. (1919) Starfishes of the Philippine seas and adjacent waters. Bulletin of the United States National Museum, 3 (100), 1–547, 156 pls.
    Fisher, W.K. (1940) Asteroidea. Discovery Reports, 20, 69–306.
    Fujita, T. & Rowe, F.W.E. (2002) Podospherasteridae fam. Nov. (Echinodermata: Asteroidea: Valvatida) with a new species, Podospheraster toyoshiomaruae from Southern Japan. Species Diversity, 7, 317–332. https://doi.org/10.12782/specdiv.7.317
    Gosliner, T.M., Behrens, D.W. & Williams, G.C. (1996) Coral Reef Animals of the Indo-Pacific. Sea Challengers, Monterey, California, 314 pp.
    Grainger, E.H. (1966) Sea Stars (Echinodermata: Asteroidea) of Arctic North America. Bulletin of the Fisheries Research Board of Canada, 152, 1–70.
    Gray, J.E. (1840) XXXII. A synopsis of the genera and species of the class Hypostoma (Asterias, Linnaeus). Annals of the Magazine of Natural History, Series 1, 6 (37), 275–290. https://doi.org/10.1080/03745484009443296
    Halpern, J.A. (1970a) Goniasteridae (Echinodermata: Asteroidea) of the Straits of Florida. Bulletin of Marine Science, 20 (1), 193–286.
    Halpern, J.A. (1970b) A monographic revision of the goniasterid sea stars of the North Atlantic. Unpublished PhD Thesis, University of Miami, Coral Gables, Florida, 253 pp.
    Hayashi, R. (1938) Sea-stars in the vicinity of the Seto Marine Biological Laboratory. Bulletin of the Biogeographical Society of Japan, 8, 271–292, pls. 5–7.
    Hayashi, R. (1938b) Sea stars of the Ogasawara Islands. Annotationes Zoological Japonenses, 17 (1), 59–68.
    Hayashi, R. (1973b) The Sea-Stars of Sagami Bay. Biological Laboratory, Imperial Household, Tokyo, 114 pp., 18 pls.
    Hernández-Agreda, A., Huckeba, J., Prata, K.E., Vermeij, M.J.A. & Bongaerts, P. (2024) Hybridization and inbreeding affect the survival of a critically endangered coral. Current Biology, 34 (21), e4, 5120–5129. https://doi.org/10.1016/j.cub.2024.09.035
    Holstein, D.M., Smith, T.B., Gyory, J. & Paris, C.B. (2015) Fertile fathoms: Deep reproductive refugia for threatened shallow corals. Scientific Reports, 5, 12407. https://doi.org/10.1038/srep12407
    Imaoka, T., Irimura, S., Okutani, T., Oguro, C., Oji, T. & Kanazawa, K. (1991) Echinoderms from the continental shelf and slope around Japan. Vol. II. Japan Fisheries Resource Conservation Association, Tokyo, 159 pp.
    iNaturalist for Pentaceraster alveolatus (2023) Available from: https://www.inaturalist.org/observations?taxon_id=255918 (accessed 22 March 2023)
    James, D.B. (1973) Studies on Indian Echinoderms. 5. New and Little Known starfishes from the Indian Seas. Journal of the Marine Biological Association of India, 15 (2), 556–559.
    Jangoux, M. (1978) Biological results of the Snellius Expedition (Echinodermata, Asteroidea). Zoologische Mededelingen, Leiden, 52, 287–300.
    Jangoux, M. (1980) Le genre Leiaster Peters (Echinodermata, Asteroidea: Ophidiasteridae). Journal of African Zoology, 94 (1–2), 87.
    Jangoux, M. (1981a) Une nouvelle espèce d’astéride bathyale des eaux de Nouvelle-Calédonie (Echinodermata, Asteroidea). Bulletin du Muséum national d’histoire naturelle, Section A: Zoologie, Biologie et Écologie Animales, 4 (3), 709–712. https://doi.org/10.5962/p.285846
    Jangoux, M. (1981b) Echinodermes: Astéroïdes. In: Forest, J. (Ed.), Résultats des campagnes MUSORSTOM: 1. Philippines (18–28 Mars 1976). Mémoires du Muséum national d’Histoire naturelle, Série A, Zoologie, 91, pp. 457–476.
    Jangoux, M. (1984) Les asterides littoraux de Nouvelle-Calédonie. Bulletin du Museum National D’Histoire Naturelle (Zoologie), Series, 6 (2), 279–293. https://doi.org/10.5962/p.285919
    Jangoux, M. (1986) Les astérides. In: Guille, A., Laboute, P. & Menou, J.L. (Eds.), Guide des étoiles de mer, oursins et autres échinodermes du lagon de Nouvelle-Caleodnie. Faune Tropicale, 25, pp. 111–153.
    Keesing, J.K., Mortimer, N., Hellmrich, L., Godoy, D., Babcock, R.C., Heyward, A., Paton, D. & Harvey, E.S. (2023) The short spined crown-of-thorns starfish Acanthaster brevispinus is a corallivore too. Coral Reefs, 42 (2), 399–404. https://doi.org/10.1007/s00338-023-02351-1
    Koehler, R. (1910) An account of the shallow-water Asteroidea. Echinoderma of the Indian Museum, 6, 1–192. https://doi.org/10.5962/bhl.title.11614
    Kogure, Y. & Kaneko, A. (2010) A solasterid asteroid Seriaster regularis (Echinodermata, Asteroidea, Velatida) found in the Okinawa Islands: The first record from Japanese waters. Bulletin of the Biogeographical Society of Japan, 65, 97–102. [in Japanese]
    Kogure, Y., Hashimoto, J. & Yamawaki, N. (2011) A large goniasterid sea star, Astrothauma euphylacteum (Echinodermata, Asteroidea) discovered at the northeastern edge of the East China Sea: a new record in Japanese waters. Biogeography, 13, 83–86.
    Lane, D.J. & Rowe, F.W.E. (2009) A new species of Asterodiscides (Echinodermata, Asteroidea, Asterodiscididae) from the tropical southwest Pacific, and the biogeography of the genus revisited. Zoosystema, 31 (3), 419–429. https://doi.org/10.5252/z2009n3a2
    Liao, Y. (1980) The echinoderms of Xisha Islands, Guangdong Province, China. 4. Asteroidea. Studia Marina Sinica, 17, 153–171, 6 pls. [in Chinese with English summary]
    Liao, Y. & Clark, A.M. (1995) The Echinoderms of Southern China. Science Press, Beijing, 614 pp., 23 pls.
    Linchangco, G.V., Foltz, D.W., Reid, R., Williams, J., Nodzak, C., Kerr, A.M., Miller, A.K., Hunter, R., Wilson, N.G., Nielsen, W.J., Mah, C.L. & Rouse, G.W., Wray, G.A. & Janies, D.A. (2017) The phylogeny of extant starfish (Asteroidea: Echinodermta) including Xyloplax based on comparative transcriptomics. Molecular Phylogenetics and Evolution, 115, 161–170. https://doi.org/10.1016/j.ympev.2017.07.022
    Livingstone, A.A. (1931) On the restriction of the genus Ferdina Gray. Australian Zoology, 6 (4), 305–309.
    Livingstone, A.A. (1934) Two new asteroids from Australia. Records of the Australian Museum, 19 (3), 177–180. https://doi.org/10.3853/j.0067-1975.19.1934.697
    Livingstone, A.A. (1936) Descriptions of new Asteroidea from the Pacific. Records of the Australian Museum, 19 (6), 383–390. https://doi.org/10.3853/j.0067-1975.19.1936.708
    Lopes, E.M., Pérez‐Portela, R., Paiva, P.C. & Ventura, C.R.R. (2016) The molecular phylogeny of the sea star Echinaster (Asteroidea: Echinasteridae) provides insights for genus taxonomy. Invertebrate Biology, 135 (3), 235–244. https://doi.org/10.1111/ivb.12135
    de Loriol, P. (1883) Catalogue Raisonne des Echinodermes recueillis par M.V. de Robillard a l’iIle Maurice. Memoires de la Société de physique et d’histoire naturelle de Genève, 28 (8), 1–64.
    Loya, Y., Puglise, K.A. & Bridge, T.C.L. (Eds.) (2019) Mesophotic Coral Ecosystems. Springer, Cham, XV + 1003 pp. https://doi.org/10.1007/978-3-319-92735-0
    Macan, T.T. (1938) Asteroidea. Scientific Reports, John Murray Expedition 1933–1934, 4 (9), 323–435, pls. 1–6.
    Mah, C. (1998) Preliminary phylogeny and taxonomic revision of the Brisingida (Asteroidea: Forcipulatacea). In: Mooi, R. & Telford, M. (Eds.), Proceedings of the 9th International Echinoderm Conference. Ballkema, San Francisco, California, pp. 273–277.
    Mah, C. (1999) Re-description and taxonomic notes on the South Pacific brisingidan Brisingaster robillardi (Asteroidea) with new ontogenetic and phylogenetic information. Zoosystema, 21 (3), 535–546. https://doi.org/10.5962/p.270149
    Mah, C.L. (2005) A phylogeny of Iconaster and Glyphodiscus (Goniasteridae; Valvatida; Asteroidea) with descriptions of four new species. Zoosystema, 27 (1), 131–167.
    Mah, C.L. (2017) Overview of the Ferdina–like Goniasteridae (Echinodermata: Asteroidea) including a new subfamily, three new genera and fourteen new species. Zootaxa, 4271 (1), 1–72. https://doi.org/10.11646/zootaxa.4271.1.1
    Mah, C.L. (2018) New genera, species and occurrence records of Goniasteridae (Asteroidea; Echinodermata) from the Indian Ocean. Zootaxa, 4539 (1), 1–116. https://doi.org/10.11646/zootaxa.4539.1.1
    Mah, C.L. (2020) New Species, Occurrence Records and Observations of Predation by Deep–sea Asteroidea (Echinodermata) from the North Atlantic by NOAA Ship Okeanos Explorer. Zootaxa, 4766 (2), 201–260. https://doi.org/10.11646/zootaxa.4766.2.1
    Mah, C.L. (2021) The East Pacific/South Pacific Boundary: New Taxa and Occurrences from Rapa Nui (Easter Island), New Caledonia and adjacent regions. Zootaxa, 4980 (3), 401–450. https://doi.org/10.11646/zootaxa.4980.3.1
    Mah, C.L. (2022) New Genera, Species and Occurrence of Deep-Sea Asteroidea (Valvatacea, Forcipulatacea, Echinodermata) collected from the Tropical Pacific Ocean by the CAPSTONE Expedition. Zootaxa, 5164 (1), 1–75. https://doi.org/10.11646/zootaxa.5164.1.1
    Mah, C.L. (2023) A new species of Astrosarkus from Western Australia including new Mesophotic occurrences of Indian Ocean Oreasteridae (Valvatida, Asteroidea). Memoirs of Museum Victoria, 82, 143–165. https://doi.org/10.24199/j.mmv.2023.82.08
    Mah, C.L. (2024) New Taxa and Occurrences of Mesophotic and Deep-sea Goniasteridae (Valvatida, Asteroidea) from Okinawa and adjacent regions. Zootaxa, 5403 (1), 1–41. https://doi.org/10.11646/zootaxa.5403.1.1
    Mah, C.L., Kogure, Y., Fujita, T. & Higashiji, T. (2024) New Taxa and Occurrences of Mesophotic and Deep-sea Goniasteridae (Valvatida, Asteroidea) from Okinawa and adjacent regions. Zootaxa, 5403 (1), 1–41. https://doi.org/10.11646/zootaxa.5403.1.1
    Mah, C. & Blake, D.B. (2012) Global Diversity and Phylogeny of the Asteroidea (Echinodermata). PLoS ONE, 7 (4), e35644. https://doi.org/10.1371/journal.pone.0035644
    Mah, C.L. & Foltz, D.W. (2011) Molecular Phylogeny of the Valvatacea (Asteroidea, Echinodermata). Zoological Journal of the Linnean Society, 161, 769–788. https://doi.org/10.1111/j.1096-3642.2010.00659.x
    Mah, C. & Fujita, T. (2020) New species and occurrence records of Japanese Solasteridae and Ganeriidae including a new species of Paralophaster from the North Pacific with an overview of Hyalinothrix. Zootaxa, 4750 (1), 67–100. https://doi.org/10.11646/zootaxa.4750.1.4
    Marsh, L.M. (1976) Western Australian Asteroidea since H.L. Clark. Thalassia Jugoslavica, 12 (1), 213–225.
    Marsh, L.M. (1977) Coral Reef Asteroids of Palau, Caroline Islands. Micronesica, 13 (2), 251–281.
    Marsh, L.M. (2000) Echinoderms of the Montebello Islands. Records of the Western Australian Museum Supplement, 59, 21–27.
    Marsh, L.M. (2009) A new species of Thromidia (Echinodermata: Asteroidea) from Western Australia. Records of the Western Australian Museum, 25, 145–151. https://doi.org/10.18195/issn.0312-3162.25(2).2009.145-151
    Marsh, L.M. & Fromont, J. (2020) Field Guide to Shallow-Water Seastars of Australia. Western Australian Museum, Welshpool, 543 pp.
    Mauzey, K.P., Birkeland, C. & Dayton, P.K. (1968) Feeding behavior of asteroids and escape responses of their prey in the Puget Sound region. Ecology, 49, 603–619. https://doi.org/10.2307/1935526
    McKnight, D.G. (2006) The marine fauna of New Zealand, Echinodermata: Asteroidea (Sea-stars). 3. Orders Velatida, Spinulosida, Forcipulatida, Brisingida with addenda to Paxillosida, Valvatida. NIWA Biodiversity Memoir, 120, 1–187.
    Meadows, P.S., Meadows, A. & Murray, J.M.H. (2012) Biological modifiers of marine benthic seascapes: Their role as ecosystem engineers. Geomorphology, 157–158, 31–48. https://doi.org/10.1016/j.geomorph.2011.07.007
    Mecho, A., Easton, E.E., Sellanes, J., Gorny, M. & Mah, C. (2019) Unexplored diversity of the mesophotic echinoderm fauna of the Easter Island ecoregion. Marine Biology, 166, 1–17. https://doi.org/10.1007/s00227-019-3537-x
    Miller, J.E. (1984) Systematics of the ophidiasterid sea stars Copidaster lymani A.H. Clark and Hacelia superba H.L. Clark (Echinodermata: Asteroidea) with a key to the species of Ophidiasteridae from the western Atlantic. Proceedings of the Biological Society of Washington, 97 (1), 194–208.
    Miller, K., Speed, C., Parnum, I., Wilson, N., Kok, J., Colquhoun, J. & Case, M. (2021) Ashmore Reef Mesophotic Reef Survey and Sampling: Voyage Report. Report prepared for Parks Australia. AIMS, Townsville, Queensland, 21 pp.
    Mortensen, T. (1925) Papers from Dr. Th. Mortensen’s Pacific Expedition 1914-16. XXIX. Echinoderms of New Zealand and the Auckland-Campbell Islands. III–V. Asteroidea, Holothuroidea and Crinoidea. Videnskabelige Meddelelser fra Dansk naturhistorisk Forening i København, 79 (15), 261–420.
    Mortensen, T. (1927) Handbook of the Echinoderms of the British Isles. Oxford University Press, London, 471 pp. https://doi.org/10.5962/bhl.title.6841
    Müller, J. & Troschel, F.H. (1840) Ueber die Gattungen der Asterien. Archiv für Naturgeschichte, 6, 318–326.
    Müller, J. & Troschel, F.H. (1842) System der Asteriden.1. Asteriae. 2. Ophiuridae. Vieweg, Braunschweig, xxx + 134 pp., 12 pls. https://doi.org/10.5962/bhl.title.11715
    Nardo, J.D. (1834) De Asteriis. Isis von Oken, 1834, 716–717.
    Nance, J.M. & Braithwaite, L.F. (1979) The function of mucous secretions in the cushion star Pteraster tesselatus Ives. Journal of Experimental Marine Biology and Ecology, 40 (3), 259–266. https://doi.org/10.1016/0022-0981(79)90055-8
    Nobili, G. (1904) Diagnoses preliminaires de vingt-huit especes nouvelles de Stomatopodes et Decapodes Macroures de la Mer Rouge. Bulletin du Muséum national d’histoire naturelle, 10, 228–238.
    O’Hara, T., Mah, C.L., Hipsley, C.A., Bribiesca-Contreras, G. & Barrett, N.S. (2018) The Derwent River seastar: re-evaluation of a critically endangered marine invertebrate. Zoological Journal of the Linnean Society, 186 (2), 483–490. https://doi.org/10.1093/zoolinnean/zly057
    O’Loughlin, P.M. & Waters, J.M. (2004) A molecular and morphological revision of genera of Asterinidae (Echinodermata: Asteroidea). Memoirs of Museum Victoria, 61, 1–40. https://doi.org/10.24199/j.mmv.2004.61.1
    Paine, R.T. (1966) Food web complexity and species diversity. American Naturalist, 100, 65–75. https://doi.org/10.1086/282400
    Paine, R.T. (1969) The Pisaster-Tegula interaction: prey patches, predator food preference and intertidal community structure. Ecology, 50, 950–961. https://doi.org/10.2307/1936888
    Perrier, E. (1875) Revision de la Stellerides du Museum d’Historie Naturelle de Paris. Archives du Zoologie Experimentale et Generale, 4, 265–450.
    Perrier, E. (1881) Report on the Results of dredging in the Gulf of Mexico and in the Caribbean Sea, 1877–79, by the United States Coastal Survey Steamer Blake. 14. Description sommaire des espèces nouvelles d’Astéries. Bulletin of the Museum of Comparative Zoology, 9, 1–31.
    Perrier, E. (1894) Stellérides. Expéditions Scientifique Travailleur et du Talisman, 3, 1–431, 26 pls.
    Peters, W. (1852) Ubersicht der Seesterne (Asteridae) von Mossambique. Bericht über die zur Bekanntmachung geeigneten Verhandlungen der Konigl.Preuss.Akademie der Wissenschaften zu Berlin, 1852, 177–178.
    Pettibone, M. (1969) remarks on the North Pacific Harmothoe tenebricosa Moore (Polychaeta, Polynoidae) and its association with asteroids (Echinodermata, Asteroidea). Proceedings of the Biological Society of Washington, 82, 31–42.
    Pope, E.C. & Rowe, F.W.E. (1977) A new genus and two new species in the family Mithrodiidae (Echinodermata: Asteroidea) with comments on the status of species of Mithrodia Gray, 1840. Australian Zoology, 19 (2), 201–216.
    Pyle, R.L., Boland, R., Bolick, H., Bowen, B.W., Bradley, C.J., Kane, C., Kosaki, R.K., Langston, R., Longenecker, K., Montgomery, A., Parrish, F.A., Popp, B.N., Rooney, J., Smith, C.M. & Wagner, D. (2016) A comprehensive investigation of mesophotic coral ecosystems in the Hawaiian Archipelago. PeerJ, 4, e2475, 1–45. https://doi.org/10.7717/peerj.2475
    Riegl, B. & Piller, W.E. (2003) Possible refugia for reefs in times of environmental stress. International Journal of Earth Science, 92, 520–531. https://doi.org/10.1007/s00531-003-0328-9
    Rowe, F.W.E. (1977) A new family of Asteroidea (Echinodermata), with the description of five new species and one new subspecies of Asterodiscides. Records of the Australian Museum, 31 (5), 187–233. https://doi.org/10.3853/j.0067-1975.31.1977.209
    Rowe, F.W.E. (1977) The status of Nardoa (Andora) A.M. Clark, 1967 (Asteroidea: Ophidiasteridae), with the description of two new subgenera and three new species. Records of the Australian Museum, 31 (6), 235–245. https://doi.org/10.3853/j.0067-1975.31.1977.210
    Rowe, F.W.E. (1985) Six new species of Asterodiscides A.M. Clark (Echinodermata, Asteroidea) with a discussion of the origin and distribution of the Asterodiscididae and other ‘amphi-Pacific echinoderms. Bulletin du Muséum national d’histoire naturelle, Section A: Zoologie, Biologie et Écologie Animales, 7 (3), 531–577. https://doi.org/10.5962/p.285862
    Rowe, F.W.E. (1985b) On the genus Podosphaeraster A.M. Clark and Wright (Echinodermata, Asteroidea) with description of a new species from the North Atlantic. Bulletin du Muséum national d’histoire naturelle, Section A, Zoologie, Biologie et Écologie Animales, Series 4, 7 (2), 309–329. https://doi.org/10.5962/p.287568
    Rowe, F.W.E. (1989) Nine new deep-water species of Echinodermata from Norfolk Island and Wanganella Bank, northeastern Tasman Sea, with a checklist of the echinoderm fauna. Proceedings of the Linnean Society of New South Wales, 111 (4), 257–291.
    Rowe, F.W.E. & Gates, J. (1995) Echinodermata. In: Wells, A. (Ed.), Zoological Catalogue of Australia. Vol. 33. CSIRO, Melbourne, pp. 1–510.
    Rowe, F.W.E. & Marsh, L.M. (1982) A revision of the asterinid genus Nepanthia Gray 1840 (Echinodermata: Asteroidea), with the description of three new species. Memoirs of the Australian Museum, 16, 89–120. https://doi.org/10.3853/j.0067-1967.16.1982.360
    Rowe, F.W.E. & Nichols, D. (1980) A new species of Podosphaeraster Clark and Wright, 1962 (Echinodermata: Asteroidea) from the Pacific. Micronesica, 16 (2), 289–295.
    Ryansky, A. (2020) Starfishes and other echinoderms of the tropical Indo-Pacific. ReefID books, Middletown, DE, 91 pp.
    Sladen, W.P. (1889) Asteroidea. Report of the Scientific Results of H.M.S. Challenger, 30, 1–893. https://doi.org/10.1111/j.1096-3642.1882.tb02281.x
    Soota, T.D. & Sastry, D.R.K. (1979) Notes on two species of Echinaster Muller & Troschel (Echinodermata: Asteroidea) from the Indian Ocean. Records of the Zoological Survey of India, 343–352. https://doi.org/10.26515/rzsi/v75/i1-4/1979/161889
    Spencer, W.K. & Wright, C.W. (1966) Asterozoans. Part U. Echinodermata. In: Moore, R.C. (Ed.), Treatise on Invertebrate Paleontology. 3 (1). University of Kansas Press, Lawrence, Kansas, pp. U4–U107.
    Verrill, A.E. (1867) Notes on Radiata in the Museum of Yale College, with descriptions of new genera and species. No. 3. On the geographical distribution of the echinoderms of the West coast of America. Transactions of the Connecticut Academy of Arts and Sciences, 1 (2), 323–351.
    Verrill, A.E. (1899) Revision of certain genera and species of starfishes, with descriptions of new forms. Transactions of the Connecticut Academy of Arts and Sciences, 10 (1), 145–234. https://doi.org/10.5962/bhl.part.7031
    Viguier, C. (1878) Classification des Stellerides. Comptes rendus hebdomadaires des séances de l’Académie des sciences, 86, 681–683.
    Wagner, R.H., Phillips, D.W., Standing, J.D. & Hand, C. (1979) Commensalism or mutualism: Attraction of a sea star towards its symbiotic polychaete. Journal of Experimental Marine Biology and Ecology, 39 (3), 205–210. https://doi.org/10.1016/0022-0981(79)90126-6
    Walenkamp, J.H.C. (1976) The asteroids of the coastal waters of Surinam. Zoologische Verhandelingen, 147, 1–91, 18 pls.
    Zhang, R., Fau, M., Mah, C., Eléaume, M., Zhang, D., Zhou, Y., Lu, B. & Wang, C. (2024) Systematics of deep-sea starfish order Brisingida (Echinodermata: Asteroidea), with a revised classification and assessments of morphological characters. Molecular Phylogenetics and Evolution, 191, 107993. https://doi.org/10.1016/j.ympev.2023.107993