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Published: 2019-04-05
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First Dysmorphoptilidae from the Permian of China (Hemiptera: Cicadomorpha: Prosbolomorpha), with notes on the fossil record of the family

Laboratory of Evolutionary Entomology and Museum of Amber Inclusions, Department of Invertebrate Zoology and Parasitology, Faculty of Biology, University of Gdańsk, 59, Wita Stwosza St., PL80-308 Gdańsk, Poland
State Key Laboratory of Palaeobiology and Stratigraphy, Center for Excellence in Life and Paleoenvironment, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China
Hemiptera Permocixiellinae subfam. nov. Linglunxiellus gen. nov. L. chaohuensis gen. and sp. nov. Eoscarterellinae stat. nov. Permian China Dysmorphoptilidae evolution palaeobiogeography

Abstract

The first representative of Dysmorphoptilidae from the Middle Permian of China is described. Linglunxiellus chaohuensis gen. and sp. nov. is placed together with Permocixiella venosa Becker-Migdisova, 1961 from Kuznetsk Basin in the newly established subfamily Permocixiellinae subfam. nov., Eoscarterellinae stat. nov. is proposed as separate subfamily within Dysmorphoptilidae. The classification and fossil record of Dysmorphoptilidae is reviewed, and an annotated catalogue of included taxa in Dysmorphotiloidea is presented.

References

  1. Adami-Rodrigues, K. & Iannuzzi, R. (2001) Late Paleozoic terrestrial arthropod faunal and floral successions in the Paraná Basin: a preliminary synthesis. Acta Geologica Leopoldensia, 52/53, 165–179.

    Adami-Rodrigues, K., Iannuzzi, R. & Pinto, I.D. (2004) Permian plant-insect interactions from a Gondwana flora of southern Brazil. Fossils and Strata, 51 (51), 106–125.

    Amyot, C. J.-B. & Audinet-Serville, J.G. (1843) Histoire naturelle des insectes Hémiptères. Paris, Libraire Encyclopédique de Roret, 675 pp.

    Aristov, D.S. (2013) 1.3. Vyatkian-Olenkian localities. In: Aristov, D.S., Bashkuev, A.S., . Golubev, V.K., Gorochov, A.V., Karasev, E.V., Kopylov, D.S., Ponomarenko, A.G., Rasnitsyn, A.P., Rasnitsyn, D.A., Sinitshenkova, N.D., Sukatsheva , I.D., Vassilenko, D.V. (2013) Fossil Insects of the Middle and Upper Permian of European Russia. Paleontological Journal, 47 (7), 641–832.

    https://doi.org/10.1134/S0031030113070010

    Beattie, R. (2007) The geological setting and palaeoenvironmental and palaeoecological reconstructions of the Upper Permian insect beds at Belmont, New South Wales, Australia. African Invertebrates, 48 (1), 41–57.

    Becker-Migdisova, E.E. [Bekker-Migdisova, E.E.] (1946) Ocherki po sravnitel’noy morfologii sovremennykh i permskikh Homoptera. Chast’ I [Contributions to the knowledge of comparative morphology of recent and Permian Homoptera. Part. I]. Izviestia AN SSSR, Biology Series, 6, 741–766 [in Russian].

    Becker-Migdisova, E.E. [Bekker-Migdisova, E.E.] (1949) Mezozoyskie Homoptera Sredney Azii. [Mesozoic Homoptera of Middle Asia]. Trudy Paleontologicheskogo Instituta AN SSSR, 22, 1–68 [in Russian].

    Becker-Migdisova, E.E. [Bekker-Migdisova, E.E.] (1955) Iskopaemye nasekomye iz triasa Sibiri. [Fossil insects from the Triassic of Siberia]. Doklady Akademii Nauk SSSR, 105, 1100–1103.

    Becker-Migdisova, E.E. [Bekker-Migdisova, E.E.] (1960) Novye permskie ravnokrylye evropeyskoy chasti SSSR. [New Permian Homoptera from the European U.S.S.R.]. Trudy Paleontologicheskogo Instituta, Akademia Nauk SSSR, 7, 1–112 [in Russian].

    Becker-Migdisova, E.E. [Bekker-Migdisova, E.E.] (1961) Rhynchota. In: Rohdendorf, B.B., Becker-Migdisova, E.E., Martynova, O.M. & Sharov, A.G. (Eds.), Paleozoic insects of the Kuznetsk Basin. Trudy Paleontologicheskogo Instituta, Akademia Nauk SSSR, 85, 286–392. [in Russian].

    Becker-Migdisova, E.E. [Bekker-Migdisova, E.E.] (1962) Otryad Homoptera. In: Orlov, Y.A. (Head-Editor), Osnovy Paleontologii, Rohdendorf, B.B. (editor of volume). Chlenistonogie, Trakheïnye i khelicerovye, 9. Izdatel’stvo Akademii Nauk SSSR, Moscow, pp. 162–208 [in Russian].

    Becker-Migdisova, E.E. [Bekker-Migdisova, E.E.] (1991) Order Homoptera. In: Rohdendorf, B.B. (Ed.), Fundamentals of Paleontology. Arthropoda, Tracheata, Chelicerata, 9, pp. 219–289. Smithsonian Institution Libraries and the National Science Foundation, Washington, D.C.

    Bourgoin, T., Wang, R., Asche, M., Hoch, H., Soulier-Perkins, A., Stroinski, A., Yap, S. & Szwedo, J. (2015) From micropterism to hyperpterism: recognition strategy and standardized homology-driven terminology of the forewing venation patterns in planthoppers (Hemiptera: Fulgoromorpha). Zoomorphology, 134, 63–77.

    https://doi.org/10.1007/s00435-014-0243-6

    Brauckmann, C., Martins-Neto, R.G. & Gallego, O.F. (2006) The Triassic insect fauna from Argentina. – Blattoptera, Glosselytrodea, Miomoptera, Auchenorrhyncha, and Coleoptera from the Los Rastros Formation (Bermejo Basin), Los Chañares locality (La Rioja Province). Clausthaler Geowissenschaften, 5, 1−9.

    Burmeister, H. (1835). Schnabelkerfe. Rhynchota. Handbuch der Entomologie, Besondere Entomologie, 2 (1), T.C.F. Enslin, Berlin, 396 pp.

    Carpenter, F.M. (1992) Hexapoda. Treatise on Invertebrate Paleontology, Part R, Arthropoda 4 (3). Geological Society of America and University of Kansas, Boulder, Colorado, and Lawrence, Kansas, xxi + 277 pp.

    Chen, J., Szwedo, J., Wang, B., Zheng, Y., Jiang, H., Jiang, T., Wang, X. & Zhang, H-C. (2019) A new bizarre cicadomorph family in mid-Cretaceous Burmese amber (Hemiptera, Clypeata). Cretaceous Research, 97, 1–15.

    https://doi.org/10.1016/j.cretres.2019.01.010

    Cohen, K.M., Finney, S.C., Gibbard, P.L. & Fan, J.-X. (2013; updated) The ICS International Chronostratigraphic Chart. Episodes, 36, 199–204.

    Devincenzi, S.M. (2018) Indicadores de crecimiento y de uso para las colecciones paleontológicas del IANIGLA-CCT CONICET Mendoza (Argentina). Revista del Museo de La Plata, 3 (2), 324–334.

    https://doi.org/10.24215/25456377e062

    Dohrn, A. (1859) Catalogus Hemipterorum. Herausgegeben von dem Entomologischen verein zu Stettin. Herrcke & Lebeling, Stettin, 112 pp.

    Duméril, C.A.-M. (1805) Zoologie analytique, ou méthode naturelle de classification des animaux, rendue plus facile à l’aide de tableaux synoptiques. Allais, Paris, xxxii + 344 pp.

    Edwards, M.A. & Tobias, M.A. (1993) Nomenclator zoologicus. Vol. 8 (1966–1977). The Zoological Society of London, London, [4] +620 pp.

    Edwards, M.A. & Vevers, H.G. (1975) Nomenclator zoologicus. Vol. 7 (1956–1965). The Zoological Society of London, London, [4] + 374 pp.

    Edwards, M.A., Manly, P. & Tobias, M.A. (1996) Nomenclator zoologicus. Vol. 8 (1978–1994). The Zoological Society of London, London, [2] + 747 pp.

    Emeljanov, A.F. (1987) Phylogeny of Cicadina (Homoptera) based on comparative morphological data. Transactions of the All-Russia Entomological Society [Trudy Vsesoyuznogo Entomologicheskogo Obshchestva], 69, 19–109 [in Russian].

    Evans, J.W. (1943) Upper Permian Homoptera from New South Wales. Records of the Australian Museum, 21 (4), 180−198.

    https://doi.org/10.3853/j.0067-1975.21.1943.534

    Evans, J.W. (1946) A natural classification of leaf-hoppers. Transactions of the Royal entomological Society of London, 96, 47−60.

    Evans, J.W. (1956) Paleozoic and Mesozoic Hemiptera (Insecta). Australian Journal of Zoology, 4, 165–268.

    https://doi.org/10.1071/ZO9560165

    Evans, J.W. (1958) New Upper Permian Homoptera from the Belmont Beds. Records of the Australian Museum, 24 (9), 109–114.

    https://doi.org/10.3853/j.0067-1975.24.1958.647

    Evans, J.W. (1959) The zoogeography of some Australian insects. In: Keast, A., Crocker, R.L., Christina, C.S. (Eds.), Biogeography and ecology in Australia. Springer Science + Business Media, B.V., Dordrecht, pp. 150–163.

    https://doi.org/10.1007/978-94-017-6295-3_9

    Evans, J.W. (1961) Some Upper Triassic Homoptera from Queensland. Memoirs of the Queensland Museum, 14, 13–23.

    Evans, J.W. (1963a) Systematic position of the Ipsviciidae (Upper Permian Hemiptera) and some new Upper Permian and Middle Triassic Hemiptera from Australia (Insecta). Journal of the Entomological Society of Queensland, 2, 17–23.

    Evans, J.W. (1963b) The phylogeny of the Homoptera. Annual Review of Entomology, 8, 77–94.

    Evans, J.W. (1971) Some Upper Triassic insects from Mt. Crosby, Queensland. Memoirs of the Queensland Museum, 16 (1), 145–162.

    Gallego, O.F. & Martins-Neto, R.G. (1999) La entomofauna mesozoica de la Argentina: Estado actual del conocimiento. Revista de la Sociedad Entomológica Argentina, 58, 86–94.

    Gallego, O.F. & Martins-Neto, R.G. (2006) Propuesta preliminar sobre la distribución estratigráfica de las faunas triásicas de insectos y conchostracos de Argentina, Chile y sur de Brasil. Universidad Nacional del Nordeste Comunicaciones Científicas y Tecnológicas 2006, B-009, 1–4.

    Gallego, O.F., Martins-Neto, R.G. & Carmona, M.J. (2001) Nuevos registros de artrópodos (Insecta y Conchostraca) en el Triásico de la Argentina: comentarios sobre su afinidad con faunas de Laurasia y Gondwana. Universidad Nacional del Nordeste (Argentina), Comunicaciones Científicas y Tecnológicas, B-057, 1–4.

    Gallego, O.F., Martins-Neto, R.G. & Nielsen, S.N. (2005) Conchostracans and insects from the Upper Triassic of the Biobío river (‘Santa Juana Formation’), south-central Chile. Revista Geológica de Chile, 32 (2), 293–311.

    https://doi.org/10.4067/S0716-02082005000200007

    Giebel, C.G. (1856) Fauna der Vorwelt mit steter Berücksichtigung der lebenden Thiere. Monographisch dargestellt. Zweiter Band: Gliederthiere, Erste Abtheilung: Insekten und Spinnen. F.A. Brockhaus, Leipzig, xviii + 511 pp.

    Hamilton, K.G.A. (1992) Lower Cretaceous Homoptera from the Koonwarra Fossil Bed in Australia with a new superfamily and synopsis of Mesozoic Homoptera. Annals of the Entomological Society of America, 85 (4), 423–430.

    https://doi.org/10.1093/aesa/85.4.423

    Handlirsch, A. (1906-1908) Die Fossilen Insekten und die Phylogenie der Rezenten Formen. Ein Handbuch für Paläontologen und Zoologen. Verlag von Wilhelm Engelmann, Leipzig, 1430 pp.+51 Pls.

    Handlirsch, A. (1921) Paläeontologie. In: Schröder, C. (Ed.), Handbuch der Entomologie, 3 (6), 209–304.

    Handlirsch, A. (1939) Neue Untersuchungen über die fossilen Insekten mit Ergänzungen und Nachträgen sowie Ausblicken auf phylogenetische, paläogeographische und allgemein biologische Probleme. II. Teil. Annalen des Naturhistorischen Museums in Wien, 49, 1−240.

    Hong, Y.C. & Chen, R. (1981) Magnacicadiidae, a new family of Homoptera from the Middle Triassic of Tongchuan, Shaanxi Province. Chinese Science Bulletin, 26 (4), 338−340.

    Huang, D.Y., Nel, A., Lin Q.B. & Dong F.B. (2007) The first Glosselytrodea (Insecta) from the latest Middle Permian of Anhui Province, China. Bulletin de la Société Entomologique de France, 112, 179−182.

    Jeannel, R. (1949) Les insectes. Classification et phylogénie. Les insectes fossiles. In: Grassé, P.-P. (Ed.), Traité de zoologie. Anatomie, systématique, biologie, 9, 110 pp.

    Jell, P.A. (2004) The fossil insects of Australia. Memoirs of the Queensland Museum, 50 (1), 1–124.

    Kukalová-Peck, J. (1991) Fossil history and the evolution of hexapod structures. In: CSIRO, (ed.) The Insects of Australia, 2nd ed., Vol. 1, 141–179, Melbourne University Press, Carlton, Victoria, pp. 141–179.

    Labandeira, C.C. (2005) The fossil record of insect extinction: new approaches and future directions. American Entomologist, 51 (1), 14–29.

    https://doi.org/10.1093/ae/51.1.14

    Labandeira, C.C. Anderson, J.M. & Anderson, H. (2018) Expansion of arthropod herbivory in Late Triassic South Africa: The Molteno Biota, Aasvoëlberg 411 site and developmental biology of a gall. In: Tanner, L.H. (Ed.), The Late Triassic World. Earth in time of transition. Topics in Geobiology 46. Springer International Publishing AG, Cham, pp. 623–719.

    https://doi.org/10.1007/978-3-319-68009-5_14

    Lambkin, K.J. (2015) Revision of the Dysmorphoptilidae with emarginate tegmina (Hemiptera: Auchenorryncha: Cicadomorpha: Prosboloidea) of the Queensland Triassic. Zootaxa, 3936 (3), 357–374.

    https://doi.org/10.11646/zootaxa.3936.3.3

    Lambkin, K.J. (2016) Revision of the Dysmorphoptilidae (Hemiptera: Cicadomorpha: Prosboloidea) of the Queensland Triassic—Part 2. Zootaxa, 4092 (2), 207–218.

    https://doi.org/10.11646/zootaxa.4092.2.4

    Lara, M.B. & Wang, B. (2016) New hemipteran insects (Eoscarterellidae, Scytinopteridae, and Protopsyllidiidae) from the Upper Triassic Potrerillos Formation of Mendoza, Argentina. Paläontologische Zeitschrift, 90, 49–61.

    https://doi.org/10.1007/s12542-016-0286-8

    Lara, M.B., Cariglino, B. & Zavattieri, A.M. (2017) Palaeoenvironmental interpretation of an Upper Triassic deposit in southwestern Gondwana (Argentina) based on an insect fauna, plant assemblage, and their interactions. Palaeogeography, Palaeoclimatology, Palaeoecology, 476, 163–180.

    https://doi.org/10.1016/j.palaeo.2017.03.029

    Lara, M.B., Gallego, O.F. & Tassi, L.V. (2012) Mesozoic coleopteran faunas from Argentina: geological context, diversity, taphonomic observations, and comparison with other fossil insect records. Psyche, 242563, 1–14.

    https://doi.org/10.1155/2012/242563

    Latreille, P.A. (1802) Histoire naturelle, générale et particuliere des Crustacés et des Insectes. T. 3. Famille naturelle des genres. F. Dufart, Paris, 467 pp.

    Latreille, P.A. (1806) Genera Crustaceorum et Insectorum secundum ordinem naturalem in familias disposita, iconibus exemplisque plurimis explicata, 1. Paris et Argentorati, bibliopolam, 302 pp.

    Latreille, P.A. (1807) Genera crustaceorum et insectorum: secundum ordinem naturalem in familias disposita, iconibus exemplisque plurimis explicata, 3. Amand Koenig. Paris, 258 pp.

    Leach, W.E. (1815) Entomology, In: Brewster, D. (Ed.), The Edinburgh Encyclopedia, Ed. 1, 9 (1), Edinburgh, pp. 57−172.

    Lin, Q.B. (1986) Early Mesozoic fossil insects from South China. Palaeontologia Sinica, New Series B, 170 (21), 1−112.

    Lin, Q.B., Nel, A. & Huang, D.Y. (2010) The first agetopanorpine mecopteroid insect from Middle Permian of China (Insecta: Mecoptera: Permochoristidae). Annales de la Société Entomologique de France, 46 (1–2), 62−66.

    https://doi.org/10.1080/00379271.2010.10697639

    Linnaeus, C. (1758) Systema Naturae per Regna tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. L. Salvii, Holmiae, 824 pp.

    Lozovsky, V.R., Minikh, M.G., Grunt, T.A., Kukhtinov, D.A., Ponomarenko, A.G. & Sukacheva, I.D. (2009a) Ufimskiy yarus vostochno-evropeyskoy shkaly: status, validnost’, korrelatsionnyï potentsial. Stratigrafiya. Geologicheskaya Korrelyatsiya, 17 (6), 46–58 [in Russian].

    Lozovsky, V.R., Minikh, M.G., Grunt, T.A., Kukhtinov, D.A., Ponomarenko, A.G. & Sukacheva I.D. (2009b) The Ufimian Stage of the East European Scale: Status, Validity, and Correlation Potential. Stratigraphy and Geological Correlation, 17 (6), 602–614.

    Martins-Neto, R.G. (2005) Estágio atual da paleoartropodologia brasileira: hexápodes, miriápodes, crustáceos (Isopoda, Decapoda, Eucrustacea e Copepoda) e quelicerados. Arquivos do Museu Nacional, Rio de Janeiro, 63 (3), 471–494.

    Martins-Neto, R.G. (2008) Insectos fósiles sudamericanos. In: Camacho, H.H., Longobucco, M.I. (Eds.), Los Invertebrados fósiles. Vol. 2, Fundación de Historia Natural Félix De Azara, Vazquez Mazzini Editores, Buenos Aires, pp. 673–680.

    Martins-Neto, R.G. & Gallego, O.F. (1999) The Triassic Insect Fauna from Argentina. I. Auchenorrhyncha, Miomoptera and Ensifera. Revista Española de Paleontología, 14, 191–202.

    Martins-Neto, R.G. & Gallego, O.F. (2001) The Triassic Insect Fauna from Argentina. IV. Glosselytrodea and complements on Auchenorrhyncha. Acta Geologica Leopoldensia, 24, 105–114.

    Martins-Neto, R.G. & Gallego, O.F. (2006) Review of Dysmorphoptilidae Handlirsch (Hemiptera: Cicadomorpha) from the Argentinean Triassic, with description of a new subfamily, and a new species. Polish Journal of Entomology, 75 (2), 185–197.

    Martins-Neto, R.G., Gallego, O.F & Melchor, R.N. (2003) The Triassic insect fauna from South America (Brazil, Argentina and Chile): a checklist (except Blattoptera and Coleoptera) and descriptions of new taxa. Acta Zoologica Cracoviensia, 46 (suppl. Fossil Insects), 229–256.

    Martins-Neto, R.G., Gallego, O.F. & Tassi, L.V. (2011) The Triassic coleopteran fauna of southern South America: Morphometric variation of elytra, paleobiogeography, and a phylogenetic approach. Gaea, Journal of Geoscience, 7 (1), 1–18.

    https://doi.org/10.4013/gaea.2011.71.01

    Martynov, A.V. (1937a) Liassovye nasekomye Shuraba i Kizil Kii [Liassic insects from Shurab and Kisyl Kiya]. Trudy Paleontologischeskogo Instituta Akademii Nauk SSSR, 7 (1), 1–232 [in Russian].

    Martynov, A.V. (1937b) Permskie iskopaemye nasekomye Kargaly i ikh otnosheniya [Permian fossil insects from Kargala and their relationships]. Trudy Paleontologischeskogo Instituta Akademii Nauk SSSR, 7 (2), 1–91 [in Russian].

    Matsumura, S. (1940) New species and genera of Cercopidae in Japan, Korea and Formosa, with a list of the known species. Journal of the Faculty of Agriculture, Hokkaido Imperial University, 45, 35–82.

    Metcalf, Z.P. & Wade, V. (1966) A Catalogue of the fossil Homoptera (Homoptera: Auchenorrhyncha). General Catalogue of the Homoptera. A Supplement to Fascicle I. Membracidae of the General Catalogue of the Hemiptera. Contribution from the Entomology Department, North Carolina Agricultural Experiment Station. Paper Nr. 2049, Raleigh, N.C. USA (North Carolina State University at Raleigh), v + 245 pp.

    Miroshnikov, L.D. & Pirozhnikov, L.P. (1956) O mestonakhozhdeniyakh iskopaemykh nasekomykh na territorii Sovetskoï Arktiki [On localities of fossil insect founds in Soviet Arctic territory]. Trudy Nauchno-Issledovatel’skogo Instituta Geologii Arktiki Ministerstva Geologii i Okhrany Nedr SSSR, Sbornik stateï po geologii Arktiki, 89 (6), 139–146. [in Russian].

    Morel, E.M. (1994) El Triásico del Cerro Cacheuta, Mendoza (Argentina). Parte I. Geología, contenido paleo-florístico y cronoestratigrafía. Ameghiniana, 31 (2), 161–176.

    Moura-Júnior, D.A. de, Scheffler, S.M. & Fernandes A.C.S. (2018) A paleoentomofauna Brasileira: cenário atual / The Brazilian fossil insects: current scenario. Anuário do Instituto de Geociências – UFRJ, 41, 142–166.

    https://doi.org/10.11137/2018_1_142_166

    Neave, S.A. 1939. Nomenclator zoologicus: a list of the names of genera and subgenera in zoology from the tenth edition of Linnaeus, 1758, to the end of 1935. Vol. 2 (D-L). Zoological Society of London, London, 1025 pp.

    Neave, S.A. 1940a. Nomenclator zoologicus: a list of the names of genera and subgenera in zoology from the tenth edition of Linnaeus, 1758, to the end of 1935. Vol. 3 (M-P). Zoological Society of London, London, 1065 pp.

    Neave, S.A. 1940b. Nomenclator zoologicus: a list of the names of genera and subgenera in zoology from the tenth edition of Linnaeus, 1758, to the end of 1935. Vol. 4 (Q-Z and supplement). Zoological Society of London, London, 756 pp.

    Nel, A., Prokop, J., Nel, P., Grandcolas, P., Huang, D., Roques, P., Guilbert, E., Dostal, O. & Szwedo, J. (2012) Traits and evolution of wing venation pattern in paraneopteran insects. Journal of Morphology, 273, 480–506.

    https://doi.org/10.1002/jmor.11036

    Nicholson, D.B., Mayhew, P.J. & Ross, A.J. (2015) Changes to the fossil record of insects through fifteen years of discovery. PLoS ONE, 10 (7), e0128554, 1–61.

    https://doi.org/10.1371/journal.pone.0128554

    Pérez, V. & Pérez, E. (2015) Los insectos (Insecta) del Mesozoico chileno. The insects (Insecta) from the chilean Mesozoic. Anales Instituto Patagonia (Chile), 43 (1), 165–170.

    https://doi.org/10.4067/S0718-686X2015000100017

    Pinto, I.D. (1972) Late Paleozoic insects and crustaceans from Paraná Basin and their bearing on chronology and continental drift. Simpósio internacional sobre os sistemas Carbonífero e Permiano na América do Sul. Anais do Academia Brasileira de Ciencias, Rio de Janeiro, 44 (Suplemento), 247–260.

    Pinto, I.D. (1990) Permian Insects from Paraná Basin, South Brazil. VI Homoptera – 3. Fulgoringruidae. Pesquisas, 17 (1–2), 3–6.

    Pinto, I.D. (1996) Paleobotanical and paleozoological age divergences in South American strata. Pesquisas em Geociências, 22 (1/2) [1995], 46–52.

    Ponomarenko, A.G., Yan, E.V. & Huang, D.Y. (2014) New beetles (Coleoptera) from the terminal Middle Permian of China. Paleontological Journal, 48 (2), 191−200.

    https://doi.org/10.1134/S0031030114010109

    Popov, Yu.A. (1980) Nadotryad Cimicidea Laicharting, 1781. Otryad Cimicina Laicharting, 1781. Poluzhestorylye, ili khobotnye [Superorder Cimicidea Laicharting, 1781. Order Cimicina Laicharting, 1781. Hemipterans, or rhynchotans]. In: Rohdendorf, B.B. & Rasnitsyn, A.P. (Eds.), Istoricheskoe razvitye klassa nasekomykh [Historical development of the class Insecta]. Trudy Paleontologicheskogo Instituta Akademii Nauk SSSR, 175, 58–69.

    Qadri, M.A.H. (1967) Phylogenetic study of Auchenorrhyncha. University Studies (Karachi), 4 (3), 1–16.

    Rafinesque, C.S. (1815) Analyse de la nature, ou tableau de l’univers et des corps Organisés. Pivately published, Palerme, 224 pp.

    Ricetti, J., Schneider, J., Iannuzzi, R. & Weinschütz, L.C. (2016) Anthracoblattina mendesi Pinto et Sedor (Blattodea, Phyloblattidae): the most completely preserved South American Palaeozoic cockroach. Revista Brasileira de Paleontologia, 19 (2), 181–194.

    https://doi.org/10.4072/rbp.2016.2.03

    Riek, E.F. (1968) Undescribed fossil insects from the Upper Permian of Belmont, New South Wales (with an appendix listing the described species). Records of the Australian Museum, 27 (15), 303–310.

    https://doi.org/10.3853/j.0067-1975.27.1968.449

    Riek, E.F. (1973) Fossil insects from the Upper Permian of Natal, South Africa. Annals of the Natal Museum, 21, 513–532.

    Riek, E.F. (1974) Upper Triassic insects from the Molteno “Formation”, South Africa. Palaeontographia Africana, 17, 19–31.

    Riek, E.F. (1976) A new collection of insects from the Upper Triassic of South Africa. Annales of the Natal Museum, 22 (3), 791−820.

    Santos, R.V. Souza, P.A., Souza de Alvarenga, C.J., Dantas, E.L., Martins Pimentel, M., Gouveia de Oliveira, C. & Medeiros de Araújo, L. (2006) Shrimp U-Pb zircon dating and palynology of bentonitic layers from the Permian Irati Formation, Paraná Basin, Brazil. Gondwana Research, 9, 456–463.

    https://doi.org/10.1016/j.gr.2005.12.001

    Schulze, F.E., Kükenthal, W. & Heider, K. (1928) Derot-Electre. Nomenclator animalium generum et subgenerum. Im Auftrage der Preuβischen Akademie der Wissenschaften zu Berlin. Vol. 2 (9), 957–1116.

    Schulze, F.E., Kükenthal, W. & Heider, K. (1931) Macrog-Microto. Nomenclator animalium generum et subgenerum. Im Auftrage der Preuβischen Akademie der Wissenschaften zu Berlin. Vol. 3 (15), 1939–2098.

    Schulze, F. E., Kükenthal, W., Heider, K. & Hesse, R. (1938) Tachymerops-Trichoec. Nomenclator animalium generum et subgenerum. Im Auftrage der Preuβischen Akademie der Wissenschaften zu Berlin, 5 (24): 3373–3532.

    Scudder, S.H. (1891) Index to the known fossil insects of the world, including myriapods and arachnids. Bulletin of the United States Geological Survey, 71, 1–744.

    Senter, P. (2008) Voices of the past: a review of Palaeozoic and Mesozoic animal sounds. Historical Biology, 20 (4), 255−287.

    https://doi.org/10.1080/08912960903033327

    Shcherbakov, D.E. (1984a) Sistema i filogeniya permskikh Cicadomorpha (Cimicida, Cicadina). Palaeontologicheskiï Zhurnal, 18 (2), 87–97. [in Russian].

    Shcherbakov, D.E. (1984b) A system and the phylogeny of Permian Cicadomorpha (Cimicida, Cicadina). Paleontological Zhurnal, 2, 89–101.

    Shcherbakov, D.E. (1988) Novye mesozoiskie ravnokryle. [New Mesozoic Homoptera], In Rosanov, A.Y. (Ed.) Novye iskopaemye bespozvonochnye Mongolii. [New fossil invertebrates from Mongolia]. Transactions of the joint Soviet–Mongolian Palaeontological Expedition, 33, 60–63. [in Russian].

    Shcherbakov, D.E. (1996) Origin and evolution of the Auchenorrhyncha as shown by the fossil record. In: Schaefer, C.W. (Ed.), Studies on Hemipteran phylogeny. Thomas Say publications in entomology, Entomological Society of America, Lanham (Maryland), 31–45.

    Shcherbakov, D.E. (2000) Permian faunas of Homoptera (Hemiptera) in relation to phytogeography and the Permo–Triassic crisis. Paleontological Journal, 34, Supplement 3, S251–S267.

    Shcherbakov, D.E. (2002) The 270 million year history of Auchenorrhyncha (Homoptera). In: Holzinger, W. (Ed.), Zikaden – Leafhoppers, Planthoppers and Cicadas (Insecta: Hemiptera: Auchenorrhyncha). Denisia, 4 [Kataloge des OÖ. Landesmuseums Linz, Neue Folge 176], 29–36.

    Shcherbakov, D.E. (2011) New and little-known families of Hemiptera Cicadomorpha from the Triassic of Central Asia−early analogs of treehoppers and planthoppers. Zootaxa, 2836, 1–26.

    https://doi.org/10.11646/zootaxa.2836.1.1

    Shcherbakov, D.E. & Popov Yu. A. (2002) Superorder Cimicidea Laicharting, 1781. Order Hemiptera Linné, 1758. The bugs, cicads, scale insects, etc. In: Rasnitsyn, A.P., Quicke D.L.J. (Eds.), History of insects. Kluwer Academic Publishers, Dordrecht, pp. 143–157.

    Spinola, M. (1839) Essai sur les Fulgorelles, sous-tribu de la tribu des Cicadaires, ordre des Rhyngotes. Annales de la Société Entomologique de France, 8, 133−337.

    Sukatcheva, I.D. & Rasnitsyn, A.P. (2004a) [Jurassic Insects (Insecta) from the Sai-Sagul Locality (Kyrgyzstan, Southern Fergana).] Paleontologicheskii Zhurnal, 2, 2004, 64–68 [in Russian].

    Sukatcheva, I.D. & Rasnitsyn, A.P. (2004b) Jurassic Insects (Insecta) from the Sai-Sagul Locality (Kyrgyzstan, Southern Fergana). Paleontological Journal, 38 (2), 182–186.

    Szwedo, J. (2002) Amber and amber inclusions of planthoppers, leafhoppers and their relatives (Hemiptera, Archaeorrhyncha et Clypaeorrhyncha). In: Holzinger, W. (Ed.), Zikaden – Leafhoppers, Planthoppers and Cicadas (Insecta: Hemiptera: Auchenorrhyncha). Denisia, 4 [Kataloge des OÖ. Landesmuseums Linz, Neue Folge 176], 37–56.

    Szwedo, J. (2018) The unity, diversity and conformity of bugs (Hemiptera) through time. Earth and Environmental Science Transactions of the Royal Society of Edinburgh, 107, 109–128.

    https://doi.org/10.1017/S175569101700038X

    Szwedo, J., Bourgoin, T. & Lefebvre, F. (2004) Fossil planthoppers (Hemiptera: Fulgoromorpha) of the World. An annotated catalogue with notes on Hemiptera classification. Studio 1, Warszawa, 199 pp.+8 Pls.

    Tillyard, R.J. (1919) Mesozoic Insects of Queensland. No. 7 Hemiptera Homoptera; with a note on the phylogeny of the suborder. Proceedings of the Linnean Society of New South Wales, 44, 857−896.

    Tillyard, R.J. (1922) Mesozoic insects of Queensland. No. 9. Orthoptera, and additions to the Protorthoptera, Odonata, Hemiptera and Plannipennia. Proceedings of the Linnean Society of New South Wales, 47, 447–470.

    Tillyard, R.J. (1923) Mesozoic insects of Queensland. No. 10. Summary of the upper Triassic insect fauna of Ipswich, Q. (With an appendix describing new Hemiptera and Planniptera). Proceedings of the Linnean Society of New South Wales, 48, 481–498.

    Tillyard, R.J. (1926) Upper Permian insects of New South Wales Part 1. Introduction and the order Hemiptera. Proceedings of the Linnean Society of New South Wales, 51, 1−30.

    Tillyard, R.J. (1936) A new Upper Triassic fossil insect bed in Queensland. Nature, 138, 719−720.

    https://doi.org/10.1038/138719b0

    van Dijk, D.E. & Geertsema, H. (1999) Permian Insects from the Beaufort Group of Natal, South Africa. Annals of the Natal Museum, 40, 137–171.

    Wang B., Szwedo, J. & Zhang, H.C. (2012) New Jurassic Cercopoidea from China and their evolutionary significance (Insecta: Hemiptera). Palaeontology, 55 (6), 1223–1243.

    Wang, B., Szwedo, J., Zhang, H.C. & Fang, Y. (2010) New Froghoppers from the Jurassic of China and Their Phylogenetic Significance (Insecta: Hemiptera: Cercopoidea). Earth Science Frontiers, 17, Special Issue, 224–225.

    Wang, Y., Ren, D., Liang, J.H., Liu, Y.S. & Wang, Z.H. (2006) The fossil Homoptera of China: A review of present knowledge. Acta Zootaxonomica Sinica, 31 (2), 294–303.

    Wang, Y., Shih, C.K., Shu, L. & Ren, D. (2010). Chapter 14. Homoptera – 17 years underground. In: Ren, D., Shih, C.K., Gao, T.P., Yao, Y.Z. & Zhao, Y.Y. (Eds.), Silent stories – insect fossil treasures from dinosaur era of the Northeastern China. Science Press, Beijing, 118–138.

    Wang, Y., Shih, C.K., Shu, L. & Ren, D. (2012). [Chapter 14. Homoptera – 17 years underground]. In: Ren, D., Shih, C.K., Gao, T.P., Yao, Y.Z. & Zhao, Y.Y. (Eds.), [Silent stories – insect fossil treasures from dinosaur era of the Northeastern China]. Science Press, Beijing, 152–172 [in Chinese].

    Westwood, J.O. (1838) The entomologist’s text book; an introduction to the natural history, structure, physiology, and classification of insects, including the Crustacea and Arachnida. W.S. Orr & Co., London, 432 pp.

    Westwood, J.O. (1845) Explanation of plates. In: Brodie, P.B., A history of the fossil insects in the secondary rocks of England. Van Voorst, London, pp. 115−128.

    Zherikhin, V.V. (1978) Razvitie i smena melovykh i kainozoïskikh faunisticheskikh kompleksov (trakheïnye i khelitserovye [Development and changes of the Cretaceous and Cenozoic faunal assemblages (Tracheata and Chelicerata)]. Trudy Paleontologicheskogo Instituta AN SSSR, 165, 1–198 [in Russian].