Abstract
With 18 genera and 29 species described, the extinct family Dysmorphoptilidae Handlirsch, 1906 is one of the most diverse members of the early cicadomorphan insects, widespread from the Middle Permian to the Upper Jurassic of Europe, Siberia, Central Asia, South China, South Africa, South America, and Australia (Shcherbakov, 1984, 2000; Martins-Neto et al., 2003; Lambkin, 2015, 2016; Lara & Wang, 2016; Szwedo & Huang, 2019).
References
Becker-Migdisova, E.E. (1961) Rhynchota. In: Rohdendorf, B.B., Becker-Migdisova, E.E., Martynova, O.M. & Sharov, A.G. (Eds), Paleozoic insects of the Kuznetsk Basin. Trudy Paleontologicheskogo Instituta, Akademia Nauk SSSR, 85, 286–392 [In Russian].
Bourgoin, T., Wang, R., Asche, M., Hoch, H., Soulier-Perkins, A., Stroinski, A., Yap, S. & Szwedo, J. (2015) From micropterism to hyperpterism: recognition strategy and standardized homology-driven terminology of the forewing venation patterns in planthoppers (Hemiptera: Fulgoromorpha). Zoomorphology, 134, 63–77.
https://doi.org/10.1007/s00435-014-0243-6
Evans, J.W. (1956) Paleozoic and Mesozoic Hemiptera (Insecta). Australian Journal of Zoology, 4, 165–268.
https://doi.org/10.1071/ZO9560165
Evans, J.W. (1958) New Upper Permian Homoptera from the Belmont Beds. Records of the Australian Museum, 24, 109–114.
https://doi.org/10.3853/j.0067-1975.24.1958.647
Handlirsch, B.A. (1906–1908) Die fossilen Insekten und die Phylogenie der rezenten Formen. Ein Handbuch für Paläontologen und Zoologen. Engelmann, Leipzig, 1430 pp.
https://doi.org/10.5962/bhl.title.34145
Hong, Y.C. & Chen, R. (1981) Magnacicadidae, a new family of Homoptera from the Middle Triassic of Tongchuan, Shaanxi Province. Chinese Science Bulletin, 26, 338−340.
Huang, D.Y., Nel, A., Lin, Q.B. & Dong, F.B. (2007) The first Glosselytrodea (Insecta) from the latest Middle Permian of Anhui Province, China. Bulletin de la Société Entomologique de France, 112, 179−182.
Huang, D.Y., Schubnel, T. & Nel, A. (2020) A new middle Permian orthopteran family questions the position of the order Titanoptera (Archaeorthoptera: Orthoptera). Journal of Systematic Palaeontology, 18, 1217−1222.
https://doi.org/10.1080/14772019.2020.1733112
Lambkin, K.J. (2015) Revision of the Dysmorphoptilidae with emarginate tegmina (Hemiptera: Auchenorrhyncha: Cicadomorpha: Prosboloidea) of the Queensland Triassic. Zootaxa, 3936 (3), 357–374.
https://doi.org/10.11646/zootaxa.3936.3.3
Lambkin, K.J. (2016) Revision of the Dysmorphoptilidae (Hemiptera: Cicadomorpha: Prosboloidea) of the Queensland Triassic—Part 2. Zootaxa, 4092 (2), 207–218.
https://doi.org/10.11646/zootaxa.4092.2.4
Lara, M.B. & Wang, B. (2016) New hemipteran insects (Eoscarterellidae, Scytinopteridae and Protopsyllidiidae) from the Upper Triassic Potrerillos Formation of Mendoza, Argentina. Paläontologische Zeitschrift, 90, 49–61.
https://doi.org/10.1007/s12542-016-0286-8
Lin, Q.B., Nel, A. & Huang, D.Y. (2010) The first agetopanorpine mecopteroid insect from Middle Permian of China (Insecta: Mecoptera: Permochoristidae). Annales de la Société Entomologique de France, 46, 62−66.
https://doi.org/10.1080/00379271.2010.10697639
Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. L. Salvii, Holmiae [= Stockholm], 824 pp.
https://doi.org/10.5962/bhl.title.542
Martins-Neto, R.G. & Gallego, O.F. (2006) Review of Dysmorphoptilidae Handlirsch (Hemiptera: Cicadomorpha) from the Argentinean Triassic, with description of a new subfamily, and a new species. Polish Journal of Entomology, 75, 185–197.
Martins-Neto, R.G., Gallego, O.F. & Melchor, R.N. (2003) The Triassic insect fauna from South America (Brazil, Argentina and Chile): a checklist (except Blattoptera and Coleoptera) and descriptions of new taxa. Acta Zoologica Cracoviensia, 46, 229–256.
Nel, A., Prokop, J., Nel, P., Grandcolas, P., Huang, D.Y., Roques, P., Guilbert, E., Dostál, O. & Szwedo, J. (2012) Traits and evolution of wing venation pattern in paraneopteran insects. Journal of Morphology, 273, 480–506.
https://doi.org/10.1002/jmor.11036
Pinto, I.D. (1990) Permian Insects from Paraná Basin, South Brazil. VI Homoptera – 3. Fulgoringruidae. Pesquisas, 17, 3–6.
https://doi.org/10.22456/1807-9806.21373
Popov, Y.A. (1980) Nadotryad Cimicidea Laicharting, 1781. Otryad Cimicina Laicharting, 1781. Poluzhestorylye, ili khobotnye [Superorder Cimicidea Laicharting, 1781. Order Cimicina Laicharting, 1781. Hemipterans, or rhynchotans]. In: Rohdendorf, B.B. & Rasnitsyn, A.P. (Eds), Istoricheskoe razvitye klassa nasekomykh [Historical development of the class Insecta]. Trudy Paleontologicheskogo Instituta Akademii Nauk SSSR, 175, 58–69.
Riek, E.F. (1973) Fossil insects from the Upper Permian of Natal, South Africa. Annals of the Natal Museum, 21, 513–532.
Shcherbakov, D.E. (1984) A system and the phylogeny of Permian Cicadomorpha (Cimicida, Cicadina). Paleontological Zhurnal, 2, 89–101.
Shcherbakov, D.E. (2000) Permian faunas of Homoptera (Hemiptera) in relation to phytogeography and the Permo-Triassic crisis. Paleontologicheskii Zhurnal, 34, 251–267.
Szwedo, J. (2018) The unity, diversity and conformity of bugs (Hemiptera) through time. Earth and Environmental Science Transactions of the Royal Society of Edinburgh, 107, 109−128.
https://doi.org/10.1017/S175569101700038X
Szwedo, J. & Huang, D.Y. (2019) First Dysmorphoptilidae from the Permian of China (Hemiptera: Cicadomorpha: Prosbolomorpha), with notes on the fossil record of the family. Palaeoentomology, 2 (2), 148–170.
https://doi.org/10.11646/palaeoentomology.2.2.6
Yao, X., Zhou, Y.Q. & Hinnov, L.A. (2015) Astronomical forcing of a Middle Permian chert sequence in Chaohu, South China. Earth and Planetary Science Letters, 422, 206–211.