Abstract
Clam shrimp (Spinicaudata) are worldwide distributed branchiopod crustaceans specialised in ephemeral freshwater habitats. The Carboniferous is an important period for the early evolution and diversification of clam shrimp. Compared with the rare and geographically confined fossil record of the Devonian, clam shrimp in the Carboniferous have a much wider geographical distribution and higher biodiversity. Over 20 genera of clam shrimp have been recorded in the Carboniferous all over the world, but they are sparse in China. To date, five records of Carboniferous clam shrimp have been reported from China (Pruvost, 1927; Zhang et al., 1976; Wang, 1987; Zheng et al., 1988; Liu & Fan, 1995; Liao et al., 2019). Among them, four species Lioestheria? mathieui Pruvost, 1927, Protomonocarina huixianensis Wang, 1987, Retrofractus lingyuanensis Liu & Fan, 1995, and Pemphilimnadiopsis cheni Liao, Shen & Huang, 2019, are found in the Pennsylvanian Benxi Formation in North China (Pruvost, 1927; Zhang et al., 1976; Wang, 1987; Liu & Fan, 1995; Liao et al., 2019).
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