Abstract
Lebanosmylus leae gen. et sp. nov., the youngest and first representative of the family Saucrosmylidae, is described from the marine lithographic Cenomanian limestone of the Konservat-Lagerstätte of Hjoula, Lebanon, on the basis of a wing compression. The paleobiogeographic implications are briefly discussed.
References
Azar, D., Maksoud, S., Huang, D.Y. & Nel, A. (2019) First Lebanese dragonflies (Insecta: Odonata, Aeshnoptera, Cavilabiata) from the Arabo-African mid-Cretaceous paleocontinent. Cretaceous Research, 93, 78–89. https://doi.org/10.1016/j.cretres.2018.08.025
Breitkreuz, L.C.V., Winterton, S.L. & Engel, M.S. (2017) Wing tracheation in Chrysopidae and other Neuropterida (Insecta): a resolution of the confusion about vein fusion. American Museum Novitates, 3890, 1–44. https://digitallibrary.amnh.org/handle/2246/6819
Engel, M.S., Winterton, S.L. & Breitkreuz, L.C.V. (2018) Phylogeny and evolution of Neuropterida: where have wings of lace taken us? Annual Review of Entomology, 63, 531–551. https://doi.org/10.1146/annurev-ento-020117-043127
Fang, H., Ren, D. & Wang, Y.J. (2015) Familial clarification of Saucrosmylidae stat. nov. and new saucrosmylids from Daohugou, China (Insecta, Neuroptera). PLoS One, 10 (10), e0141048. https://doi.org/10.1371/journal.pone.0141048
Fang, H., Ren, D., Liu, J.X. & Wang, Y.J. (2018) Revision of the lacewing genus Laccosmylus with two new species from the Middle Jurassic of China (Insecta, Neuroptera, Saucrosmylidae). ZooKeys, 790, 115–126. https://doi.org/10.3897/zookeys.790.28286
Heads, S.W., Martill, D.M. & Loveridge, R.F. (2005) An exceptionally preserved antlion (Insecta, Neuroptera) with colour pattern preservation from the Cretaceous of Brazil. Palaeontology, 48, 1409–1417. https://doi.org/10.1111/j.1475-4983.2005.00514.x
Jouault, C., Legendre, F., Perrichot, V., Condamine, F.L. & Nel, A. (2022) Redescription of Litholingia rhora Ren, 2002 (Neuroptera: Grammolingiidae) from the Middle Jurassic of Daohugou. Palaeoentomology, 5 (1), 20–26. https://doi.org/10.11646/palaeoentomology.5.1.3
Khramov, A.V. & Vasilenko, D.V. (2018) New records of Grammolingiidae, Saucrosmylidae, and Panfiloviidae (Insecta: Neuroptera) from the Jurassic of Mongolia and Kyrgyzstan. Paleontological Journal, 52, 1391–1400. https://doi.org/10.1134/S0031030118120109
Liu, Q., Zhang, H.C., Wang, B., Fang, Y., Zheng, D.R., Zhang, Q. & Jarzembowski, E.A. (2013) A new genus of Saucrosmylinae (Insecta, Neuroptera) from the Middle Jurassic of Daohugou, Inner Mongolia, China. Zootaxa, 3736 (4), 387–391. https://doi.org/10.11646/zootaxa.3736.4.6
Liu, Q., Zhang, H.C., Wang, B., Fang, Y., Zheng, D.R., Zhang, Q. & Jarzembowski, E.A. (2014) A new saucrosmylid lacewing (Insecta, Neuroptera) from the Middle Jurassic of Daohugou, Inner Mongolia, China. Alcheringa: An Australasian Journal of Palaeontology, 38, 301–304. https://doi.org/10.1080/03115518.2014.886849
Maksoud, S. & Azar, D. (2021) Hjoula: A remarkable mid-Cenomanian Lebanese fossil fish Lagerstätte now also promising for fossil insects. Palaeoentomology, 4 (3), 223–227. https://doi.org/10.11646/palaeoentomology.4.3.8
Menon, F. & Makarkin, V.N. (2008) New fossil lacewings and antlions (Insecta, Neuroptera) from the Lower Cretaceous Crato Formation of Brazil. Palaeontology, 51, 149–162. https://doi.org/10.1111/j.1475-4983.2007.00740.x
Ponomarenko, A.G. (1985) Setchatokrylye iz yury Sibiri i Zapadnoi Mongolii [Neuroptera from the Mesozoic of Siberia and Mongolia]. In: Rasnitsyn, A.P. (Ed.), Yuskie nasekomye Sibiri i Mongolii [Fossil Insects from Siberia and Mongolia]. Trudy Paleontologicheskogo Instituta Akademii Nauk SSSR, 211, 91–96.
Ren, D. & Yin, J. (2003) New ‘osmylid-like’ fossil Neuroptera from the Middle Jurassic of Inner Mongolia, China. Journal of the New York Entomological Society, 111, 1–11. https://www.jstor.org/stable/25010442
Shi, C.F., Béthoux, O., Shih, C.K. & Ren, D. (2012) Guyiling jianboni gen. et sp. n., an antlion-like lacewing, illuminating homologies and transformations in Neuroptera wing venation. Systematic Entomology, 37, 617–631. https://doi.org/10.1111/j.1365-3113.2012.00633.x
Shi, C.F., Winterton, S.L. & Ren, D. (2015) Phylogeny of split-footed lacewings (Neuroptera, Nymphidae), with descriptions of new Cretaceous fossil species from China. Cladistics, 31, 455–490. https://doi.org/10.1111/cla.12104
Stange, L.A. (2004) A systematic catalog, bibliography and classification of the world antlions (Neuroptera: Myrmeleontidae). Memoirs of the American Entomological Institute, 74, 1–565.
Wang, Y.J., Liu, Z.Q., Wang, X., Shih, C.K., Zhao, Y.Y., Engel, M.S. & Ren, D. (2010) Ancient pinnate leaf mimesis among lacewings. Proceedings of the National Academy of Science of the United States of America, 107, 16212–16215. https://doi.org/10.1073/pnas.1006460107
Winterton, S.L., Martins, C.C., Makarkin, V., Ardila-Camacho, A. & Wang, Y.J. (2019) Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera. Zootaxa, 4581 (1), 1–99. https://doi.org/10.11646/zootaxa.4581.1.1
Yang, Q., Makarkin, V.N. & Ren, D. (2013) A new genus of the family Panfiloviidae (Insecta, Neuroptera) from the Middle Jurassic of China. Palaeontology, 56, 49–59. https://doi.org/10.1111/j.1475-4983.2012.01157.x