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Published: 2022-08-30
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Palaeozoic insects from China with discussion on their ages

State Key Laboratory of Palaeobiology and Stratigraphy, Center for Excellence in Life and Paleoenvironment, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China
State Key Laboratory of Palaeobiology and Stratigraphy, Center for Excellence in Life and Paleoenvironment, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China
State Key Laboratory of Palaeobiology and Stratigraphy, Center for Excellence in Life and Paleoenvironment, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China
State Key Laboratory of Palaeobiology and Stratigraphy, Center for Excellence in Life and Paleoenvironment, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China
State Key Laboratory of Palaeobiology and Stratigraphy, Center for Excellence in Life and Paleoenvironment, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China
Late Palaeozoic insect fossils Carboniferous Permian

Abstract

The late Palaeozoic is the key period for understanding the origin and early evolution of insects, but Palaeozoic insects from China are poorly explored. Up to now, a total of 53 Carboniferous species, 27 Permian species and 1 Carboniferous–Permian species (12 orders or 2 superorders) have been described. Recently, with the development of biostratigraphy and isotope chronology, new insights into the stratigraphy of China and its timescale have been greatly improved. In this study, we discuss the ages of published Palaeozoic insects from China and summarize 23 Carboniferous species, 57 Permian species and 2 Carboniferous–Permian species.

References

  1. Béthoux, O., Cui, Y.Y., Kondratieff, B., Stark, B. & Ren, D. (2011) At last, a Pennsylvanian stem-stonefly (Plecoptera) discovered. BMC Evolutionary Biology, 11 (1), 1–12. https://doi.org/10.1186/1471-2148-11-248
  2. Béthoux, O., Gu, J.J. & Ren, D. (2012a) A new Upper Carboniferous stem—orthopteran (Insecta) from Ningxia (China). Insect Science, 19 (2), 153–158. https://doi.org/10.1111/j.1744-7917.2011.01468.x
  3. Béthoux, O., Gu, J.J., Yue, Y.L. & Ren, D. (2012b) Miamia maimai n. sp., a new Pennsylvanian stem—orthopteran insect, and a case study on the application of cladotypic nomenclature. Fossil Record, 15 (2), 103–113. https://doi.org/10.1002/mmng.201200008
  4. Brauckmann, C. & Schneider, J. (1996) A Lower Carboniferous insect from the Bitterfeld/Delitzsch area (Pterygota, Arnsbergian, Germany). Neues Jahrbuch für Geologie und Palaontologie, Monatshefte, 17–30. https://doi.org/10.1127/njgpm/1996/1996/17
  5. Brauckmann, C., Brauckmann, B. & Gröning, E. (1994) The stratigraphical position of the oldest known Pterygota (Insecta. Carboniferous, Namurian). Annales de la Société Géologique de Belgique, 117 (1), 47–56.
  6. Brauckmann, C., Schöllmann, L. & Sippel, W. (2003) Die fossilen Insekten, Spinnentiere und Eurypteriden von Hagen-Vorhalle. Geologie und Paläontologie in Westfalen, 59, 1–89. [In German]
  7. Bureau of Geological and Mineral Resources of Guizhou Province (1997) Stratigraphy (Lithostratic) of Guizhou Province. China University of Geoscience Press, Beijing, 306 pp. [In Chinese]
  8. Bureau of Geological and Mineral Resources of Henan Province (1997) Stratigraphy (Lithostratic) of Henan Province. China University of Geoscience Press, Beijing, 299 pp. [In Chinese]
  9. Bureau of Geological and Mineral Resources of Ningxia Hui Autonomous Region (1996) Stratigraphy (Lithostratic) of Ningxia Hui Autonomous Region. China University of Geoscience Press, Beijing, 132 pp. [In Chinese]
  10. Bureau of Geological and Mineral Resources of Shanxi Province (1997) Stratigraphy (Lithostratic) of Shanxi Province. China University of Geoscience Press, Beijing, 353 pp. [In Chinese]
  11. Bureau of Geological and Mineral Resources of Yunnan Province (1996) Stratigraphy (Lithostratic) of Yunnan Province. China University of Geoscience Press, Beijing, 366 pp. [In Chinese]
  12. Bureau of Geology and Mineral Resources of Anhui Province (1987) Regional Geology of Anhui Province. Geology Press, Beijing, 721 pp. [In Chinese]
  13. Bureau of Geology and Mineral Resources of Guizhou Province (1987) Regional Geology of Guizhou Province. Geology Press, Beijing, pp. 227–277. [In Chinese]
  14. Bureau of Geology and Mineral Resources of Yunnan Province (1990) Regional Geology of Yunnan Province. Geology Press, Beijing, pp. 160–197. [In Chinese]
  15. Chen, H.D., Hou,, Z.J., Tian, J.C., Liu, W.J. & Zhang, J.Q. (2001) Study on sequence stratigraphy of deposits and tectono-sedimentary evolution in Ordos Basin during Late Palaeozoic. Journal of Mineralogy and Petrology, 21, 16–24.
  16. Chen, L., Gu, J.J., Yang, Q., Ren, D., Blanke, A. & Béthoux, O. (2021) Ovipositor and mouthparts in a fossil insect support a novel ecological role for early orthopterans in 300 million years old forests. eLife, 10, e71006. https://doi.org/10.7554/eLife.71006
  17. Chen, R., Wang, F., Li, Z., Evans, N.J. & Chen, H.D. (2022) Detrital zircon geochronology of the Permian Lower Shihezi Formation, northern Ordos Basin, China: time constraints for closing of the Palaeo-Asian Ocean. Geological Magazine, 1–20. https://doi.org/10.1017/S0016756822000498
  18. Dai, S.F., Tian, L.W., Chou, C.L., Zhou, Y.P., Zhang, M.Q., Zhao, L., Wang, J., Yang, Z., Gao, H.Z. & Ren, D. (2008) Mineralogical and compositional characteristics of Late Permian coals from an area of high lung cancer rate in Xuan Wei, Yunnan, China: Occurrence and origin of quartz and chamosite. International Journal of Coal Geology, 76 (4), 318–327. https://doi.org/10.1016/j.coal.2008.09.001
  19. Du, S., Béthoux, O., Gu, J.J. & Ren, D. (2017) Protomiamia yangi gen. et sp. nov. (Early Pennsylvanian; Xiaheyan, China), a sexually dimorphic Palaeozoic stem-Orthoptera. Journal of Systematic Palaeontology, 15 (3), 193–204. https://doi.org/10.1080/14772019.2016.1154899
  20. Feng, G.D. & Shang, G.L. (1980) Benxi Group in the middle of the Weibei Coalfield in Shaanxi Province. Coal Geology & Exploration, (1), 18–22. [In Chinese]
  21. Fu, Y.Z. & Huang, D.Y. (2020) Sinomorphoptila incompleta gen. et sp. nov., a new dysmorphoptilid from the Middle Permian of China (Hemiptera, Cicadomorpha, Prosbolomorpha). Palaeoentomology, 3 (4), 361–365. https://doi.org/10.11646/palaeoentomology.3.4.7
  22. Gu, J.J., Béthoux, O. & Ren, D. (2011) Longzhua loculata n. gen. n. sp., one of the most completely documented Pennsylvanian Archaeorthoptera (Insecta; Ningxia, China). Journal of Paleontology, 85 (2), 303–314. https://doi.org/10.1666/10-085.1
  23. Gu, J.J., Béthoux, O. & Ren, D. (2014a) A new cnemidolestodean stem-orthopteran insect from the Late Carboniferous of China. Acta Palaeontologica Polonica, 59 (3), 689–696 [In Chinese].
  24. Gu, J.J., Béthoux, O. & Ren, D. (2017) A new, rare and distinctive species of Panorthoptera (Insecta, Archaeorthoptera) from the Upper Carboniferous of Xiaheyan (Ningxia, China). Fossil Record, 20 (2), 253–257. https://doi.org/10.5194/fr-20-253-2017
  25. Gu, J.J., Yue, Y.L., Wen, W.C., Zong, L.Y. & Ren, D. (2014b) A review of researches on Palaeozoic insects in China. Acta Entomologica Sinica, 57 (1), 123–132.
  26. Hong, Y.C. (1980) New findings of Late Palaeozoic insects from Shanxi. Geological Review, 26 (2), 89–95. [In Chinese]
  27. Hong, Y.C. (1983) Two new fossil genera of Paraplecoptera (Insecta) in China. Acta Palaeontologica Sinica, 22 (3), 359–362. [In Chinese]
  28. Hong, Y.C. (1985a) New fossil genera and species of Shanxi Formation in Xishan of Taiyuan. Entomotaxonomia, 7 (2), 83–91. [In Chinese]
  29. Hong, Y.C. (1985b) Palaeontological atlas of North China. Vol. 1. Paleozoic. Geological Publishing House, Beijing, pp. 492–510. [In Chinese]
  30. Hong, Y.C. (1998) Establishment of fossil entomofaunas and their evolutionary succession in North China. Insect Science, 5 (4), 283-300. https://doi.org/10.1111/j.1744-7917.1998.tb00324.x
  31. Huang, D.Y., Fu, Y.Z., Lian, X.N. & Nel, A. (2020a) Sinaspidoneura magnifica nov. gen., nov. sp., first Chinese Caloneurodea (Insecta: Archaeorthoptera). Geobios, 63, 33–37. https://doi.org/10.1016/j.geobios.2020.10.003
  32. Huang, D.Y., Nel, A., Lin, Q.B. & Dong, F.B. (2007) The first Glosselytrodea (Insecta) from the latest Middle Permian of Anhui Province, China. Bulletin de la Société Entomologique de France, 112 (2), 179–182.
  33. https://doi.org/10.3406/bsef.2007.16418
  34. Huang, D.Y., Schubnel, T. & Nel, A. (2020b) A new middle Permian orthopteran family questions the position of the Order Titanoptera (Archaeorthoptera: Orthoptera). Journal of Systematic Palaeontology, 18 (15), 1217–1222. https://doi.org/10.1080/14772019.2020.1733112
  35. Huang, D.Y., Su, Y.T., Li, X.R. & Wang, X.D. (2018) A new Dictyopteran fossil from the Carboniferous–Permian of Xishan (Beijing) with discussions on the geological ages of insect fossils from Beijing. Acta Palaeontologica Sinica, 57 (4), 474–480. [In Chinese] https://doi.org/10.1007/s11430-017-9253-7
  36. Huang, D.Y., Wei, G., Wei, T., Wu, Y., Wang, X.D. & Li, X.R. (2022) New dictyopteran fossils from the Pennsylvanian of West Beijing, China. Palaeoentomology, 5 (2), 161–166. https://doi.org/10.11646/palaeoentomology.5.2.9
  37. Huang, Y.Q., Zhang, P., Zhang, J.C., Tang, X., Liu, C.W. & Yang, J.W. (2020) Fractal characteristics of pores in the Longtan shales of Guizhou, southwest China. Geofluids, 2020. https://doi.org/10.1155/2020/8834758
  38. Li, Y., Wang, X.D., Hu, K.Y., Huang, X., Zhang, S.C., Zhang, B., Qi, Y.P., Chen, J.T. & Yao, L. (2021) Lithostratigraphic subdivision and correlation of the Carboniferous in China. Journal of Stratigraphy, 45 (3), 303–318. [In Chinese]
  39. Li, Y.J., Béthoux, O., Pang, H. & Ren, D. (2013a) Early Pennsylvanian Odonatoptera from the Xiaheyan locality (Ningxia, China): new material, taxa, and perspectives. Fossil Record, 16 (1), 117–139. https://doi.org/10.1002/mmng.201300006
  40. Li, Y.J., Ren, D., Pecharová, M. & Prokop, J. (2013b) A new palaeodictyopterid (Insecta: Palaeodictyoptera: Spilapteridae) from the Upper Carboniferous of China supports a close relationship between insect faunas of Quilianshian (northern China) and Laurussia. Alcheringa: An Australasian Journal of Palaeontology, 37 (4), 487–495. https://doi.org/10.1080/03115518.2013.793024
  41. Liao, H.Y., Lian, X.N., Gao, J., Cai, C.Y., Feng, Z. & Huang, D.Y. (2020) Discovery of Pemphilimnadiopsis cheni (Branchiopoda: Diplostraca: Spinicaudata) from the Benxi Formation in Taiyuan, North China and its stratigraphic significance. Palaeoentomology, 3 (6), 578–581. https://doi.org/10.11646/palaeoentomology.3.6.9
  42. Liao, H.Y., Shen, Y.B., Gao, J. & Huang, D.Y. (2019) Pemphilimnadiopsis cheni sp. nov. (Branchiopoda: Diplostraca: Spinicaudata) from the Upper Carboniferous of East Hebei, China and its biostratigraphic significance. Palaeoentomology, 2 (4), 381–389. https://doi.org/10.11646/palaeoentomology.2.4.12
  43. Lin, Q.B. & Han, W.J. (1985) A new cockroach from the Upper Shihhotse Formation (Upper Permian) in Yu County, Henan Province. Acta Palaeontologica Sinica, 24 (1), 122–125. [In Chinese]
  44. Lin, Q.B. & Liang X.Y. (1988) A Permian cockroach tegmen from Gongxian, Henan, China. Acta Palaeontologica Sinica, 27 (5), 640–642. [In Chinese]
  45. Lin, Q.B. (1978) Upper Permian and Triassic fossil insects of Guizhou. Acta Palaeontologica Sinica, 17 (3), 313–317. [In Chinese]
  46. Lin, Q.B. (1982) Atlas of palaeontology from East Region of China. Vol. 2. Later Paleozoic. Geological Publishing House, Beijing, pp. 329–332. [In Chinese]
  47. Lin, Q.B., Nel, A. & Huang, D.Y. (2010) The first agetopanorpine mecopteroid insect from Middle Permian of China (Insecta: Mecoptera: Permochoristidae). Annales de la Société Entomologique de France, 46 (1-2), 62–66. https://doi.org/10.1080/00379271.2010.10697639
  48. Liu, G.C., Zhang, H., Cai, C.Y., Yuan, A.H., Zheng, Y.Q. & Huang, D.Y. (2021) The bristletail genus Dasyleptus (Insecta: Archaeognatha: Dasyleptidae) from the Upper Permian-Lower Triassic Kayitou Formation of SouthWestern China. Historical Biology, 33 (8), 1292–1296. https://doi.org/10.1080/08912963.2019.1692342
  49. Lu, L.W., Fang, X.C., Ji, S.A. & Pang, Q.Q. (2002) A contribution to the knowledge of the Namurian in Ningxia. Acta Geoscientia Sinica, 23 (2), 165–168. [in Chinese]
  50. Ma, S.L., Li, J.H., Wang, L. Gao, H. & Zhang, J.G. (2021) Provenance, tectonic setting and source-area paleoweathering of the Upper Paleozoic sandstones in the northwestern Ordos Basin, China: evidence from whole-rock geochemistry. Carbonates Evaporites, 36, 64. https://doi.org/10.1007/s13146-021-00729-2
  51. Mathieu, F.F. (1939) La stratigraphie du basin houiller de Kaiping (Chine). Musée Royal d’Histoire Naturelle de Belgique, Bruxelles, 48 pp.
  52. Misof, B., Liu, S.L., Meusemann, K., Peters, R.S., Donath, A., Mayer, C., Frandsen, P.B., Ware, J., Flouri, T., Beutel, R.G., Niehuis, O., Petersen, M., Izquierdo-Carrasco, F., Wappler, T., Rust, J., Aberer, A.J., Aspöck, U., Aspöck, H., Bartel, D., Blanke, A., Berger, S., Böhm, A., Buckley, T.R., Calcott, B., Chen, J.Q., Friedrich, F., Fukui, M., Fujita, M., Greve, C., Grobe, P., Gu, S.C., Huang, Y., Jermiin, L.S., Kawahara, A.Y., Krogmann, L., Kubiak, M., Lanfear, R., Letsch, H., Li, Y.Y., Li, Z.Y., Li, J.G., Lu, H.R., Machida, R., Mashimo, Y., Kapli, P., McKenna, D.D., Meng, G.L., Nakagaki, Y., Navarrete-Heredia, J.L., Ott, M., Ou, Y.X., Pass, G., Podsiadlowski, L., Pohl, H., Reumont, B.M. von, Schütte, K., Sekiya, K., Shimizu, S., Ślipinśki, A., Stamatakis, A., Song, W.H., Su, X., Szucsich, N.U., Tan, M.H., Tan, X.M., Tang, M., Tang, J.B., Timelthaler, G., Tomizuka, S., Trautwein, M., Tong, X.L., Uchifune, T., Walzl, M.G., Wiegmann, B.M., Wilbrandt, J., Wipfler, B., Wong, T.K.F., Wu, Q., Wu, G.X., Xie, Y.L., Yang, S.Z., Yang, Q., Yeates, D.K., Yoshizawa, K., Zhang, Q., Zhang, R., Zhang, W.W., Zhang, Y.H., Zhao, J., Zhou, C., Zhou, L.L., Ziesmann, T., Zou, S.J., Li, Y.R., Xu, X., Zhang, Y., Yang, H.M., Wang, J., Wang, J., Kjer, K.M. & Zhou, X. (2014) Phylogenomics resolves the timing and pattern of insect evolution. Science, 346, 763–767. https://doi.org/10.1126/science.1257570
  53. Montañez, I.P. & Poulsen, C.J. (2013) The Late Paleozoic ice age: an evolving paradigm. Annual Review of Earth and Planetary Sciences, 41, 629–656. https://doi.org/10.1146/annurev.earth.031208.100118
  54. Nel, A., Roques, P., Nel, P., Prokin, A.A., Bourgoin, T., Prokop, J., Szwedo, J., Azar, D., Desutter-Grandcolas, L., Wappler, T., Garrouste, R., Coty, D., Huang, D.Y., Engel, M.S. & Kirejtshuk, A.G. (2013) The earliest known holometabolous insects. Nature, 503 (7475), 257–261. https://doi.org/10.1038/nature12629
  55. Pecharová, M., Ren, D. & Prokop, J. (2015) A new palaeodictyopteroid (Megasecoptera: Brodiopteridae) from the Early Pennsylvanian of northern China reveals unique morphological traits and intra-specific variability. Alcheringa: An Australasian Journal of Palaeontology, 39 (2), 236–249. https://doi.org/10.1080/03115518.2015.993299
  56. Peng, D.C., Hong, Y.C. & Zhang, Z.J. (2005) Namurian insects (Diaphanopterodea) from Qilianshan Mountains, China. Geological Bulletin of China, 24 (3), 219–234.
  57. Ponomarenko, A.G., Yan, E.V. & Huang, D.Y. (2014) New beetles (Coleoptera) from the terminal Middle Permian of China. Paleontological Journal, 48 (2), 191–200. https://doi.org/10.1134/S0031030114010109
  58. Scholze, F., Shen, S.Z., Backer, M., Wei, H.B., Huebner, M., Cui, Y.Y., Feng, Z. & Schneider, J.W. (2020) Reinvestigation of conchostracans (Crustacea: Branchiopoda) from the Permian–Triassic transition in southwest China. Palaeoworld, 29 (2), 368–390. https://doi.org/10.1016/j.palwor.2019.04.007
  59. Shen, B.H., Shen, S.Z., Hou, Z.S., Wu, Q., Zhang, S.C., Zhang, B., Zhang, Y.C. & Yuan, D.X. (2021) Lithostratigraphy subdivision and correlation of the Permian in China. Journal of Stratigraphy, 45 (3), 319–339. [In Chinese]
  60. Shen, B.H., Shen, S.Z., Wu, Q., Zhang, S.C., Zhang, B., Wang, X.D., Hou, Z.S., Yuan, D.X., Zhang, Y.C., Liu, F., Liu, J., Zhang, H., Shi, Y.K., Wang, J. & Feng, Z. (2022) Carboniferous and Permian integrative stratigraphy and timescale of North China Block. Science China Earth Sciences, 65 (6), 983–1011. https://doi.org/10.1007/s11430-021-9909-9
  61. Shen, S.Z., Zhang, H., Zhang, Y.C., Yuan, D.X., Chen, B., He, W.D., Mu, L., Wang, W.Q., Chen, J., Wu, Q., Cao, C.Q., Wang, Y. & Wang, X.D. (2019) Permian integrative stratigraphy and timescale of China. Science China Earth Sciences, 62 (1), 154–188. https://doi.org/10.1007/s11430-017-9228-4
  62. Su, Y., Zhang, Z.J. & Hong, Y.C. (2012) Two new ancient griffenflies (Insecta: Odonatoptera) from the Namurian of Ningxia, China. Insect Systematics & Evolution, 43 (1), 1–10. https://doi.org/10.1163/187631212X624205
  63. Sze, H.C. & Lee, H.H. (1945) Palaeozoic plants from Ninghsia. Bulletin of the Geological Society of China, 25, 227–260. https://doi.org/10.1111/j.1755-6724.1945.mp25001010.x
  64. Szwedo, J. & Huang, D.Y. (2019) First Dysmorphoptilidae from the Permian of China (Hemiptera: Cicadomorpha: Prosbolomorpha), with notes on the fossil record of the family. Palaeoentomology, 2 (2), 148–170. https://doi.org/10.11646/palaeoentomology.2.2.6
  65. Tianjin Institute of Geology and Mineral Resources (1985) Palaeontological atlas of North China. Vol. 1. Paleozoic. Geological Publishing House, Beijing, 639 pp. [In Chinese].
  66. Wang, S. (2001) On Kayitou Formation. Journal of Stratigraphy, 25 (2), 129–134.
  67. Wang, X.D., Hu, K.Y., Qie, W.K., Sheng, Q.Y., Chen, B., Lin, W., Yao, L., Wang, Q.L., Qi, Y.P., Chen, J.T., Liao, Z.T. & Song, J.J. (2019) Carboniferous integrative stratigraphy and timescale of China. Science China Earth Sciences, 62 (1), 135–153. https://doi.org/10.1007/s11430-017-9253-7
  68. Wei, D.D., Béthoux, O., Guo, Y.X., Schneider, J.W. & Ren, D. (2013) New data on the singularly rare ‘cockroachoids’ from Xiaheyan (Pennsylvanian; Ningxia, China). Alcheringa: An Australasian Journal of Palaeontology, 37 (4), 547–557. https://doi.org/10.1080/03115518.2013.808863
  69. Yao, X., Zhou, Y.Q. & Hinnov, L.A. (2015) Astronomical forcing of a Middle Permian chert sequence in Chaohu, South China. Earth and Planetary Science Letters, 422, 206–221. https://doi.org/10.1016/j.epsl.2015.04.017
  70. Zhang, B.L., Yao, S.P., Hu, W.X., Ding, H., Liu, B. & Ren, Y.L. (2019) Development of a high-productivity and anoxic-euxinic condition during the late Guadalupian in the Lower Yangtze region: Implications for the mid-Capitanian extinction event. Palaeogeography, Palaeoclimatology, Palaeoecology, 531, 108630. https://doi.org/10.1016/j.palaeo.2018.01.021
  71. Zhang, H., Cao, C.Q., Liu, X.L., Mu, L., Zheng, Q.F., Liu, F., Xiang, L., Liu, L.J. & Shen, S.Z. (2016) The terrestrial end-Permian mass extinction in South China. Palaeogeography, Palaeoclimatology, Palaeoecology, 448, 108–124. https://doi.org/10.1016/j.palaeo.2015.07.002
  72. Zhang, Z.J., Hong, Y.C., Lu, L.W., Fang, X.S. & Jin, Y.G. (2006) Shenzhousia qilianshanensis gen. et sp. nov. (Protodonata, Meganeuridae), a giant dragonfly from the Upper Carboniferous of China. Progress in Natural Science, 16 (3), 328–330.
  73. Zhang, Z.J., Schneider, J.W. & Hong, Y.C. (2013) The most ancient roach (Blattodea): a new genus and species from the earliest Late Carboniferous (Namurian) of China, with a discussion of the phylomorphogeny of early blattids. Journal of Systematic Palaeontology, 11 (1), 27–40. https://doi.org/10.1080/14772019.2011.634443
  74. Zhao, W.B., Wen, Z.G., Zhang, H., Wu, C.J., Liu, Y., Song, H.X., Zhang, L.W., Xi, Y.Y. & Sun, L. (2021) Integrated assessment of marine-continental transitional facies shale gas of the Carboniferous Benxi Formation in the Eastern Ordos Basin. Energies, 14 (24), 8500. https://doi.org/10.3390/en1424850